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Drosophila melanogaster histone deacetylase 4, transcript variant I


LOCUS       NM_001272580            6559 bp    mRNA    linear   INV 26-DEC-2023
            (HDAC4), mRNA.
ACCESSION   NM_001272580
VERSION     NM_001272580.2
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 6559)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 6559)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 6559)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 6559)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 6559)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 6559)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 6559)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 6559)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 6559)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 6559)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 6559)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 6559)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 6559)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 6559)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 6559)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 6559)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 6559)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 6559)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            On Jul 15, 2014 this sequence version replaced NM_001272580.1.
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..6559
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..6559
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /note="Histone deacetylase 4"
                     /map="11E8-11E9"
                     /db_xref="FLYBASE:FBgn0041210"
                     /db_xref="GeneID:32278"
     CDS             849..4466
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /EC_number="3.5.1.98"
                     /note="CG1770 gene product from transcript CG1770-RI;
                     CG1770-PI; HDAC4-PI; histone deacetylase 4"
                     /codon_start=1
                     /product="histone deacetylase 4, isoform I"
                     /protein_id="NP_001259509.1"
                     /db_xref="FLYBASE:FBpp0305525"
                     /db_xref="GeneID:32278"
                     /db_xref="FLYBASE:FBgn0041210"
                     /translation="MELKKSQELQKQRLINSFQEQSKQMELEHKLQLEHKYQFAVNSH
                     GAFQELRNESMVTAAAAAVAQEQHRQQLHQQQQQHQQQQQQQQHQQQQQQQQARGRDG
                     MKLKQNCSANASPEVKQILNCFILSRKSQAAASNGTTTTSPYRNRGVVKSSSGESLPA
                     GTVTSAHPYKIPQPPPSLLKYESDFPLRKTASEPNLLKIRLKQSVIERKARIGGPAGA
                     RRHERLLQAAQRRQQKNSVLTNCNSTPDSGPNSPPSAAALAVGVVGSRGSPTSAPIQE
                     ENEEGSQYQPGQRSSINDLPLFSSPSLPNISLGRPHLPNSAQAHAQVNAQVAAQAQAQ
                     AQAQAQAHAMFAALAAAQGGCGQPGYYNPLGMAFVGRQPAPLAMIPATGIAPQQPSPV
                     VRSASATSTSSSQASLVGDVAPPQAHAASTILPSSSSYMQQLGSVAGSGVNLHAAAVA
                     AAAAAAAAAGSLPPTNSHGHGHGSHAHPHPHAHGHGHGHGHGIYAGHQHNVPITDAQV
                     AQVHLHKQGHRPLGRTQSAPLPLGHPMLTGAVQLNVVQTHYENSEAERQAYEHQVVNQ
                     KVRQTVLTRSGAAAAAAAAAGVSVVREAQLKEEDDDSAAEVMDLTDKKKPPKTVLTST
                     IATSTSQNLPEALAAAAAAAAYRAPHNASSNSASATKSGIKLRDQEYLQQQREQLLLL
                     QQEEELAKSLMRPLSRTLSSPLVPLGPHGLSQIPDTGQQPAPIATSSSADHIPPVNLS
                     LPHRQHRQLMSTLYASQLRNHQPSASGSPPHKVTTGLAYDPLMLKHSCICGDNAQHPE
                     HSGRLQSVWARLNETDLVKRCDRLRARKATQEELQTVHTEAHAMLFGSNQCQLSRPKL
                     ENTLSASFVRLSCGGLGVDLDTTWNEHHTATAARMAAGCVIDLALKTAKGDLRNGFAV
                     VRPPGHHAEANLAMGFCFFNSIAIAAKLLRQRMPEVRRILIVDWDVHHGNGTQQAFYQ
                     SPDILYLSIHRHDDGNFFPGTGGPTECGSGAGLGFNVNISWSGALNPPLGDAEYIAAF
                     RTVVMPIARSFNPDIVLVSSGFDAATGHPAPLGGYHVSPACFGFMTRELLQLANGKVV
                     LALEGGYDLAAICDSAQECVRALLGDPAAPIAKAELERPPCQNAINTLQKTIAIQQTH
                     WPCVRMLEHTVGLSALETLKVEHDESETINAMAGLSMQSMHRTLSRDDSEEPMDQDET
                     K"
     misc_feature    855..>998
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /note="Glutamine-rich N-terminal helical domain of various
                     Class IIa histone deacetylases (HDAC4, HDAC5 and HDCA9);
                     Region: ClassIIa_HDAC_Gln-rich-N; cl25407"
                     /db_xref="CDD:421006"
     misc_feature    order(858..860,870..872,879..881,891..893,900..902,
                     912..917,924..929,936..938,948..950)
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /note="dimer interface A, D [polypeptide binding]; other
                     site"
                     /db_xref="CDD:197200"
     misc_feature    3165..4289
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /note="Histone deacetylases, class IIa; Region:
                     HDAC_classIIa; cd11681"
                     /db_xref="CDD:212544"
     misc_feature    order(3606..3611,3633..3638,3723..3725,3729..3731,
                     4005..4007,4125..4127)
                     /gene="HDAC4"
                     /locus_tag="Dmel_CG1770"
                     /gene_synonym="CG1770; dHDAC4; DHDAC4; Dmel\CG1770;
                     dmHDA405; GC1770; HDAC; hdac4; Hdac4; HDAC4a"
                     /note="active site"
                     /db_xref="CDD:212544"
ORIGIN      
        1 agatatcgat gtggttccag ctctaatagt cgtggatttc ggaacattaa tttgtatttt
       61 ttgcgtcgcg tcgccccaga aagtccaatc caacagcaaa tcgcgcgttt tccaaagaac
      121 aacaaccccg aaaaattcga aaataataat tttcttgttt tttttttctg ttttcctgcg
      181 gcaaaaattc caattttttt cgtaacatct aaacaacaac aacaaaaaaa gtgcgtgcgc
      241 gtgcagaagc gaaaaaaacc gaataaattg tgggcgaatc aacaagcaaa agagaatgag
      301 cgatgatgag gagaagaggc aaaaaaagca gaggcagttg tcgtagtact agtaccaaaa
      361 agaaactgcg aaaactgcta aagtgacaaa aacaacaaca gcaagtaacg aaagagaaaa
      421 gcaaaagaag gtaaaaagaa tgaaggcgaa taaaaaataa gcgcaaaaat atacgtctac
      481 aaaaaagcag cagcaacaac tacacaaaaa ttaacttaaa aaagtcgtga attcatcgtt
      541 cagtatatat atatatatat gtatatatat atataaatca acacacacac atacgaagag
      601 tgagagcgag agagcgtgcg ttatttattt tatattttcg ttaatcattt ttcgcatttt
      661 gtgaaacaag ttggaaaagc gacaatacaa aatacagcaa caacaatagc agcagtaagc
      721 agtttgcacc acttggaaga gatctgccat cagaagccgg aagcgatgag cgattgctgc
      781 acataacgcc ggccactctg acattggact tcaagcccca tcccgccgtg gatatcgatc
      841 agcagatcat ggagctcaag aagagccagg agctgcaaaa gcagcggctt atcaactcgt
      901 ttcaggagca atcgaagcaa atggaattgg agcataaact tcaattggag cacaagtatc
      961 aatttgcggt gaattcacat ggcgctttcc aggaattgcg aaacgagagc atggtgacag
     1021 ccgccgctgc tgcggtggct caggagcaac atcgccaaca gttgcatcaa cagcagcagc
     1081 aacatcagca gcagcagcag cagcagcaac atcaacagca gcagcagcag caacaggcac
     1141 gcggcaggga tggcatgaaa ctcaagcaga actgtagtgc caatgccagt cccgaggtca
     1201 aacagattct caactgcttc atcctgagca gaaagtccca ggcggcggca tcgaatggca
     1261 caacgacaac gtcgccctac aggaatcgcg gcgtggtgaa gagctcctcg ggcgaatccc
     1321 tcccagctgg aaccgtgacc agtgcgcatc cgtacaaaat acctcagccg ccaccctcac
     1381 tgctcaaata tgaatctgat tttccgctga ggaagacagc atccgaaccg aacctgctga
     1441 agatccggct gaagcagagc gtcatcgagc gcaaggcccg catcggagga ccggcgggag
     1501 cccggcgcca cgaacgactc ctccaggcgg cgcagcgtag gcagcaaaag aactcggttc
     1561 tcacaaactg caacagtaca ccggattctg gacccaattc accgccctcg gcagctgcgt
     1621 tggcggtggg cgtggtcggt agccgtggat caccgacaag tgcacccatt caggaggaaa
     1681 acgaggaggg cagccaatat cagccgggcc agaggagcag catcaacgat ttgccattgt
     1741 tcagttcacc atcgctgccg aatatttcgc tcgggcgacc gcatttaccc aactcggcgc
     1801 aggcgcatgc ccaggtgaat gcccaggttg ctgcccaggc tcaggcacag gcccaggcac
     1861 aggcacaggc gcatgccatg ttcgcggcac tggctgccgc ccagggtggc tgcggacagc
     1921 cgggctacta taatccactg ggcatggcat tcgttggccg gcaaccagca ccgctggcca
     1981 tgattccggc cacggggata gcaccgcagc aaccgtcgcc ggtggtgcgc agtgcatcgg
     2041 ccacttcgac atcgtcgtcg caggcctcgc tggtgggcga tgtggcgcca ccgcaggctc
     2101 atgccgcctc caccattctg ccctcgtcat cgtcctacat gcagcaactg ggcagtgtgg
     2161 ccggttcggg tgttaatctc catgccgccg ccgttgctgc agcggcagca gcagcagccg
     2221 ccgccggttc actgccaccg accaatagcc atggtcacgg acacggttcc cacgcacatc
     2281 cacatccaca tgcccatgga cacgggcatg gccatggcca tggtatctac gctggccacc
     2341 agcacaatgt gcccataacg gacgcccagg tggcgcaggt gcatctgcac aaacagggcc
     2401 atcggccgct gggacgaacg cagtcggcac cactgcccct tggacatccg atgctaaccg
     2461 gagctgtgca actgaatgtg gtccaaacgc actacgagaa tagtgaggcg gagcgccagg
     2521 cgtacgagca ccaggtggtg aaccagaaag tccgccagac cgtcctgact cgcagcggag
     2581 ctgctgcagc agccgctgct gccgccggcg tgagcgtggt gcgtgaggca cagttgaagg
     2641 aggaggatga cgactcggcc gccgaggtga tggacctcac agataagaaa aaaccgccga
     2701 agacggtgct aaccagcacg atagccacca gtacgtccca gaatctgccc gaagctttgg
     2761 cggcggcggc ggcggcagcc gcctaccgtg ccccgcacaa cgcgtccagt aactccgcct
     2821 ccgccacaaa gtccggtatt aagctgcggg accaggagta tttgcaacag cagcgggagc
     2881 aactattgct ccttcagcag gaggaggaac tggcgaagag cctaatgcgt ccgctatcgc
     2941 gaacgcttag cagtccgctg gtgccgctgg ggccacatgg tcttagtcag attcccgaca
     3001 ctggacaaca gccggcaccg atagccacat cctcgtcggc cgatcatata ccgcccgtta
     3061 acctctcgct gccgcatcgc cagcaccgcc agctaatgag cacactatac gccagccaat
     3121 tgcgtaacca ccagccatcg gcgagtggtt caccgccgca caaggtcacc accggtttgg
     3181 cctacgatcc gcttatgctg aagcattcgt gtatttgcgg agacaatgcc cagcatccgg
     3241 agcacagtgg tcgactgcag agcgtgtggg cacggttaaa tgaaacggat ctggtgaagc
     3301 gttgcgatcg cctgcgcgct cgcaaggcga cacaggagga gctgcagact gtgcacaccg
     3361 aagcgcatgc catgctcttc ggttcgaatc agtgccagct cagcaggccc aagttggaaa
     3421 acacgttgtc ggccagcttt gtgcgtttgt cgtgcggcgg cttgggtgtg gatctggata
     3481 ccacgtggaa tgagcaccat acggcaaccg ctgcacgaat ggcagccggt tgtgttatcg
     3541 atttggcact gaagacggcc aagggtgacc tgcggaatgg ctttgccgtt gtccggccgc
     3601 cgggccatca tgcggaggcc aatttggcca tgggcttttg tttcttcaat tcgatagcca
     3661 ttgcggccaa gctgctgcgt cagcggatgc ccgaggtgcg gcgcatcctg atcgtcgatt
     3721 gggatgtgca tcacggcaat ggcacacagc aagcattcta ccaaagtccc gacattctat
     3781 atctttccat acatcgacac gatgacggta acttctttcc cggcacaggt ggacccacag
     3841 agtgcggctc cggtgctggt ctcggcttta acgtgaacat ctcatggtct ggggcactta
     3901 atccgccact gggcgacgcc gagtatatcg ctgcattccg taccgttgtg atgcccatcg
     3961 cgcggagctt taatccggac attgtgctgg tatcctccgg cttcgatgcg gccaccggcc
     4021 atccggcacc gctgggtggc taccatgtct ctccggcctg ctttgggttc atgacccgcg
     4081 aactccttca gttggccaac ggcaaagtgg tgctggccct cgagggcggc tacgatctgg
     4141 ccgccatctg tgattccgcc caggagtgtg tgcgggcgct gctcggcgat cccgctgctc
     4201 ccattgccaa ggccgagctg gagcgcccgc cttgccagaa tgccatcaat acgctccaga
     4261 agacgatagc catacagcaa acgcattggc cctgcgtgag gatgctggag cacacggttg
     4321 gcttgtctgc gctggagacg ctcaaggtgg agcacgacga gtccgagacg atcaacgcca
     4381 tggctggcct ctcgatgcag tcgatgcaca gaactctatc ccgcgatgat tccgaggagc
     4441 cgatggatca ggatgaaacc aaatagaagg cgatgcagac gtatattgcg gaagaggagg
     4501 cggtgggcgt ggcgaggttt ttgtggacct ggtagagtgg gaatggcagg agggggcgtg
     4561 gccgcaagct gctcaatgca catttagtta tttgttgtag ttgaattgat tgttgttgct
     4621 gttgttgttg tggcgctcgc ttgcaattgt tgtcgttgtc attgatcgtc tatagccaca
     4681 tctatctctc tcactcgatc gattatagca tcccaattcc ccgccccgcc aaaaaaaaca
     4741 aaaagaagag aaacatcgaa aatatagggg aacatgtcta aaagaagcag aacagggcct
     4801 aaagttcaaa atggtggctg tgcctccaaa agaagaaata attataataa ttagaaaaga
     4861 atcctttcaa aaaaaaagga gattttatat gaatatatat atatatgaat atatatgtag
     4921 cttaagctca gaatttagtt gtagaagtca tcgagatttt ttttttttgt tttaaggtga
     4981 gagcagaggt attttaacgc acacacaaac acacgcaact caatgtatct tattattatt
     5041 atttaagtaa gtgtgtcaaa aggatataga accactaacg tactggtgaa catcatcagt
     5101 aagctataaa gataatgttt aaagaggtcg ggtgcaagaa gcatgtgttt ttttttataa
     5161 ggtaaggttc aacctcaaag ttcccaagtc tagaccccac cacgcccact tatccgccat
     5221 cgcttacaga gatcgagtgg ggagttgatt acatgttatg tagaatgctt tagttgataa
     5281 taagatatga gaaaaaaaaa acttaacaac taactagttg gatgaaaagc gaaaaaactg
     5341 agaaatcttt ttgtacaatt tatagtttag tgcaagtttc aagtacaaca tgctattatt
     5401 taaagaattc tatatatata catatatata tatatatata caaaaaaaat atatattgtg
     5461 caataaatgt gagaaaatga aaatcgccac acaaaacaga aactaaaacg aataagagaa
     5521 aatttgttgt taaatgttgc cgagatctta tatacgtatg tatacctaat aactgttaca
     5581 ataataaccc tttatatata taactatata aatgcacatg tgtattcacc atgtgtgtaa
     5641 cgataacgat gttcaacaaa tgtctgaaat gaagtaaaac tattattata tgtaaataaa
     5701 acacaaaaaa caaagagaac aattttaata aaaaaaaagc aacaataaaa tacaaaatca
     5761 aagaaaatat ccacgctcac acacacacat acacacacaa acatatatac gtcctgggca
     5821 ggacccgaga ggcgtgaatc gggaaaaatt ctgaaataac cgttcaatcg ctaaccgtta
     5881 actgtaaaac cgctgtaact gtatttgtac tcgtgtacct ttagctgtct atagccgtct
     5941 ctctctaact cgctacctaa ctgtaaagga gcacctcaat aacgaaagaa gaaggagcaa
     6001 cagacaagga tcaaaaatat atagcaagtg aaaaggatga tggaaagaag gaggaggagg
     6061 aggagaaaca agaaaaataa gcacaacgag agatggaccg attggctcca actggatgaa
     6121 acaattgtta acaaaacgaa acagaaacta aaaactatcc ttaaaaaaaa atgaaatcac
     6181 aaatgtaact attaatctaa gcgaagcata tgcccttgcc atttttattt tcttattatt
     6241 cacctgtttt ctcttttcaa tacagcagat tttttgaaaa acatactttt gaacataagc
     6301 tgtttcgtat gcaagccacg taattgctgt tacttaggaa tatatatata cgaatttttc
     6361 ttgggttcca aaagcgttac aatacgctct gtgttaaatc tgtgctttta ttttaatttt
     6421 ctatttataa tttggcaaat tggcgatctc cccttacctt accttttttt atctttcctg
     6481 aatttgtttt ttttttacca aaacacgaat tatgaagagg attctactaa caagatgaaa
     6541 tattattgtt gcgcactgc