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Drosophila melanogaster Tousled-like kinase (Tlk), transcript


LOCUS       NM_001272293            7490 bp    mRNA    linear   INV 26-DEC-2023
            variant M, mRNA.
ACCESSION   NM_001272293
VERSION     NM_001272293.1
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 7490)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 7490)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 7490)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 7490)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 7490)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 7490)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 7490)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 7490)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 7490)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 7490)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 7490)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 7490)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 7490)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 7490)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 7490)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 7490)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 7490)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 7490)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..7490
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..7490
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Tousled-like kinase"
                     /map="3E6-3F1"
                     /db_xref="FLYBASE:FBgn0283657"
                     /db_xref="GeneID:318206"
     CDS             396..4124
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /EC_number="2.7.11.1"
                     /note="CG34412 gene product from transcript CG34412-RM;
                     CG34412-PM; Tlk-PM; BEST:GH17330; tousled-like kinase;
                     tousled like kinase"
                     /codon_start=1
                     /product="Tousled-like kinase, isoform M"
                     /protein_id="NP_001259222.1"
                     /db_xref="FLYBASE:FBpp0305895"
                     /db_xref="GeneID:318206"
                     /db_xref="FLYBASE:FBgn0283657"
                     /translation="MSPGAHLQMSPQNTSSLSQHHPHQQQQLQPPQQQQQHFPNHHSA
                     QQQSQQQQQQEQQNPQQQAQQQQQILPHQHLQHLHKHPHQLQLHQQQQQQLHQQQQQH
                     FHQQSLQGLHQGSSNPDSNMSTGSSHSEKDVNDMLSGGAATPGAAAAAIQQQHPAFAP
                     TLGMQQPPPPPPQHSNNGGEMGYLSAGTTTTTSVLTVGKPRTPAERKRKRKMPPCATS
                     ADEAGSGGGSGGAGATVVNNSSLKGKSLAFRDMPKVNMSLNLGDRLGGSAGSGVGAGG
                     AGSGGGGAGSGSGSGGGKSARLMLPVSDNKKINDYFNKQQTGVGVGVPGGAGGNTAGL
                     RGSHTGGGSKSPSSAQQQQTAAQQQGSGVATGGSAGGSAGNQVQVQTSSAYALYPPAS
                     PQTQTSQQQQQQQPGSDFHYVNSSKAQQQQQRQQQQTSNQMVPPHVVVGLGGHPLSLA
                     SIQQQTPLSQQQQQQQQQQQQQQLGPPTTSTASVVPTHPHQLGSLGVVGMVGVGVGVG
                     VGVNVGVGPPLPPPPPMAMPAAIITYSKATQTEVSLHELQEREAEHESGKVKLDEMTR
                     LSDEQKSQIVGNQKTIDQHKCHIAKCIDVVKKLLKEKSSIEKKEARQKCMQNRLRLGQ
                     FVTQRVGATFQENWTDGYAFQELSRRQEEITAEREEIDRQKKQLMKKRPAESGRKRNN
                     NSNQNNQQQQQQQHQQQQQQQNSNSNDSTQLTSGVVTGPGSDRVSVSVDSGLGGNNAG
                     AIGGGTVGGGVGGGGVGGGGVGGGGGRGLSRSNSTQANQAQLLHNGGGGSGGNVGNSG
                     GVGDRLSDRGGGGGGIGGNDSGSCSDSGTFLKPDPVSGAYTAQEYYEYDEILKLRQNA
                     LKKEDADLQLEMEKLERERNLHIRELKRILNEDQSRFNNHPVLNDRYLLLMLLGKGGF
                     SEVHKAFDLKEQRYVACKVHQLNKDWKEDKKANYIKHALREYNIHKALDHPRVVKLYD
                     VFEIDANSFCTVLEYCDGHDLDFYLKQHKTIPEREARSIIMQVVSALKYLNEIKPPVI
                     HYDLKPGNILLTEGNVCGEIKITDFGLSKVMDDENYNPDHGMDLTSQGAGTYWYLPPE
                     CFVVGKNPPKISSKVDVWSVGVIFYQCLYGKKPFGHNQSQATILEENTILKATEVQFS
                     NKPTVSNEAKSFIRGCLAYRKEDRMDVFALARHEYIQPPIPKHGRGSLNQQQQAQQQQ
                     QQQQQQQQQQSSTSQANSTGQTSFSAHMFGNMNQSSSS"
     misc_feature    <2001..>2411
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Family of unknown function (DUF5401); Region:
                     DUF5401; pfam17380"
                     /db_xref="CDD:375164"
     misc_feature    3135..3938
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Catalytic domain of the Serine/Threonine kinase,
                     Tousled-Like Kinase; Region: STKc_TLK; cd13990"
                     /db_xref="CDD:270892"
     misc_feature    order(3144..3146,3183..3185,3189..3191,3297..3299,
                     3348..3359,3369..3371,3375..3377,3492..3494,3498..3500,
                     3504..3509,3513..3515,3555..3557,3564..3566,3633..3644)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="active site"
                     /db_xref="CDD:270892"
     misc_feature    order(3369..3371,3375..3377,3492..3494,3498..3500,
                     3504..3506,3564..3566,3633..3644)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="polypeptide substrate binding site [polypeptide
                     binding]; other site"
                     /db_xref="CDD:270892"
     misc_feature    order(3552..3590,3603..3644)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="activation loop (A-loop); other site"
                     /db_xref="CDD:270892"
ORIGIN      
        1 gaactttgac attgtaatgt tttcactgag gatttgatag ggtattcaac ttttcgtctt
       61 ttgctttcaa tccgtcattc gtgagttttt attttgtgcg aattttctgt tctgtcgcct
      121 tttattatta ttactacagt tttatatatt tttttgtgtg aagcgtactc tgccgacgtc
      181 gctgccactg ccagcgctgt cgctttgacg atttgcgctt tgcgttgcgc ttactcacaa
      241 tcaaatctca cgccgcattt ttagtctcaa taataactga aacgaaaaac acgcgcagcc
      301 gaaacgaagc gcaaaccaaa gagtaaagcg aaagaagcaa ggaaggcaaa cggcgaacaa
      361 aaacgcaaaa cgaaaacgac gacacgtcgt caaacatgtc ccccggcgcc catttgcaga
      421 tgtccccgca gaacacttcg tccctaagtc aacaccatcc acatcaacag caacagttac
      481 aacccccaca gcagcaacaa cagcatttcc ctaaccatca cagcgcccag caacagtcgc
      541 agcagcagca gcaacaggag caacagaatc cccagcagca ggcgcaacag cagcagcaga
      601 tactcccaca tcaacatttg cagcacctgc acaagcatcc gcatcagctg caactgcatc
      661 agcagcagca acaacaactc caccagcaac agcagcaaca cttccaccag cagtcgctgc
      721 aagggctgca tcagggtagc agcaatccgg attcgaatat gagcactggc tcctcgcata
      781 gcgagaagga tgtcaatgat atgctgagtg gcggtgcagc aacgccagga gctgcagcag
      841 cagcgattca acagcaacat cccgcctttg cgcccacact gggaatgcag caaccaccgc
      901 cgcccccacc tcaacactcc aataatggag gcgagatggg ctacttgtcg gcaggcacga
      961 ccacgacgac gtcggtgtta acggtaggca agcctcggac gccagcggag cggaaacgga
     1021 agcgaaaaat gcctccatgt gccactagtg cggatgaggc ggggagtggc ggtggctctg
     1081 gcggagcagg agcaaccgtt gttaacaaca gcagcctgaa gggcaaatca ttggcctttc
     1141 gtgatatgcc caaggtaaac atgagcctga atctgggcga tcgtctggga ggatctgcag
     1201 gaagcggagt aggagccggt ggcgccggaa gcgggggagg tggcgctggt tccggttctg
     1261 gaagcggtgg cggcaaaagc gcccgcctga tgctgccagt cagcgacaac aagaagatca
     1321 acgactattt caataagcag caaacgggcg tgggcgtcgg tgtgccaggt ggtgcgggag
     1381 gcaataccgc tggccttcga ggatcacata cgggaggtgg cagcaagtca ccctcatccg
     1441 cccagcagca gcaaacggcg gcacagcagc agggaagcgg tgttgcgacg ggaggcagtg
     1501 caggcggttc cgctggcaac caggtgcaag tgcaaacgag cagcgcttac gccctttacc
     1561 caccagctag tccccaaacc cagacgtcac agcaacagca gcagcagcaa ccgggatcag
     1621 actttcacta tgtcaactcc agcaaggcgc agcaacaaca gcagcgtcaa cagcaacaga
     1681 cttccaatca aatggttcct ccacacgtgg tcgttggcct tggtggtcat ccactgagcc
     1741 tcgcgtccat tcagcagcag acgcccttat cccagcagca acagcagcaa caacagcagc
     1801 agcaacagca gcaactggga ccaccgacca catcgacggc ctccgtcgtg ccaacgcatc
     1861 cgcatcaact cggatccctg ggagttgttg ggatggtcgg tgtgggtgtt ggcgtgggcg
     1921 ttggagtaaa tgtgggtgtg ggaccaccac tgccaccacc accgccgatg gccatgccag
     1981 cggccattat cacttatagt aaggccactc aaacggaggt gtcgctgcat gaattgcagg
     2041 agcgcgaagc ggagcacgaa tcgggcaagg tgaagctaga cgagatgaca cggctgtccg
     2101 atgaacaaaa gtcccaaatt gttggcaacc agaagacgat tgaccagcac aagtgccaca
     2161 tagccaagtg tattgatgtg gtcaagaagc tgttgaagga gaagagcagc atcgagaaga
     2221 aggaggcgcg acagaagtgc atgcagaatc gcctcaggct cggacagttt gttacccaac
     2281 gagtgggcgc cacattccag gagaactgga cggacggcta tgcgttccag gagctgagtc
     2341 ggcggcaaga agaaataacc gctgagcgtg aagagataga tcggcagaaa aagcagctga
     2401 tgaaaaagcg tccggcggag tccggacgca agcgcaacaa caacagtaac cagaacaacc
     2461 agcagcagca gcaacagcaa caccagcaac agcagcagca acaaaattcc aactcgaacg
     2521 attccacgca gctgacgagc ggagttgtta ccggtccagg cagtgatcgt gtgagcgtaa
     2581 gcgtcgacag cggattgggt ggcaataatg cgggcgcgat cggtggcgga accgttggtg
     2641 gtggcgttgg aggtggtggt gttggaggcg gtggtgtcgg aggcggcggt ggacgtggac
     2701 tttctcgcag caattcgacg caggccaatc aggctcaatt gctgcacaac ggcggtggtg
     2761 gttcgggcgg caatgtcggc aactcgggcg gcgttggcga ccgcttgtca gatcgaggag
     2821 gaggaggtgg cggcatcggc ggaaacgata gcggcagctg ctcggactcg ggcactttcc
     2881 tgaagccaga ccccgtatcg ggtgcctaca cagcgcagga gtattacgag tacgatgaga
     2941 tcctcaagtt gcgacaaaat gccctcaaaa aggaggacgc cgacctgcag ctggagatgg
     3001 agaagctgga gcgggagcgc aatctgcaca tccgagagct caagcggatt cttaacgagg
     3061 atcagtcccg ctttaacaat catcccgtgc tgaatgatcg ctatcttctg ttgatgctcc
     3121 tgggcaaggg cggcttctca gaggtccaca aggccttcga cctgaaggag caacgctatg
     3181 tcgcatgtaa ggtgcaccaa ttaaacaagg attggaagga ggataagaaa gctaattata
     3241 tcaaacacgc tttgcgggaa tacaacattc acaaggcact ggatcatccg cgggtcgtca
     3301 agctatacga tgtcttcgag atcgatgcga attccttttg cacagtgctc gaatactgtg
     3361 atggccacga tctggacttc tatttgaagc aacataagac tatacccgag cgtgaagcgc
     3421 gctcgataat aatgcaggtt gtatctgcac tcaagtatct aaatgagatt aagcctccag
     3481 ttatccacta cgatctgaag cccggcaaca ttctgcttac cgagggcaac gtctgcggcg
     3541 agattaagat caccgacttc ggtctgtcaa aggtgatgga cgacgagaat tacaatcccg
     3601 atcacggcat ggatctgacc tctcaggggg cgggaaccta ctggtatctg ccacccgagt
     3661 gctttgtcgt gggcaaaaat ccgccgaaaa tctcctccaa agtggacgta tggagtgtgg
     3721 gtgttatctt ctaccagtgt ctgtacggca aaaagccctt cggtcacaat cagtcgcagg
     3781 ccacgattct cgaggagaat acgatcctga aggccaccga agtgcagttc tccaacaagc
     3841 caaccgtttc taacgaggcc aagagtttca ttcggggatg cttggcctat cgcaaggagg
     3901 atcgcatgga tgtgttcgca ctggccaggc acgagtacat tcagccaccg ataccgaaac
     3961 atgggcgcgg ttcgctcaat cagcaacagc aggcgcaaca acagcagcag caacaacagc
     4021 aacagcagca gcaacagtcg tcgacgtcac aggccaattc tacaggccag acatctttct
     4081 ctgcccacat gtttggcaat atgaatcagt cgagttcgtc ctagctgcca accaagcgca
     4141 attcgaactc ataattcgcc gccgccgcag tagcagcagc tacattagct gccagtagtc
     4201 aacagcaact tcaacaccag caacagcaac tccaaatcca gcaataccag cagaatcagc
     4261 aacttcattt gcaaaacagc agtagcaatt atgcggccac gcttctagat gcagcggcct
     4321 tttgctacga aaccgtgatg gatgcagccc acttccaaaa caacaataat aacagcacca
     4381 tcatcaagaa caatatgcag acaatgccgg caaacgctgc caatgctgct gcagctgcag
     4441 ggggaggggg aacaccgacc tatcagggga aatacaagcg ctggcagcag caacagttac
     4501 aacagcaggc gcaactccaa cagcagcagc acaagctgct gttgcagcaa gagctgctct
     4561 aaaatatgaa gggaatgatg acaagcgagt ttaagagaac gaggagctgc ctgccacact
     4621 gccgccgtca gcaataacaa catcccaaga aaattcacag cgagtcgcaa aagcatcaac
     4681 aaacaaagca aaaaacgcaa aaaaggatta caaagacgca gagaataggg gacatttgca
     4741 atgggaaata acaaattatt gcattatctg ctggattgtg attgtggttc aaacattaaa
     4801 aatacactta ctcatacaca caaggtagac actgcagact aattttgcct taaaagaaac
     4861 aaaacgaata atgttaactg taaattttca ctttgtgtcc atgcatagtt actagtaatg
     4921 atgatataaa tatataaata tatataaaat ataaaaatta tataaatgta taaatttata
     4981 tttatatata tatatatata tatatatata tatctgcata aacattgaag catactagta
     5041 aatctttata aatatattta tatatatatt tatctcttcc ttttaacgct tttaatcgga
     5101 aaaaacatcc caattatatt ttaattcaat ttgaaatata aaagtcattt ttgtttgttt
     5161 tgtatcaaaa tccaaaaaaa attgatttaa atataaaaat accataaaca ttaacaccac
     5221 acacacacac acacacacat atatatatat atatatatat atatatatat atacatatat
     5281 atatacattt gtatatagtc gtatctgaac cacaaacaaa tccacagata aaaaaaactg
     5341 aaaaattcat agaatcaaac cattttttta tatgccacaa agaaaagaaa ataagtagaa
     5401 atgcaagaaa aatcgagaaa aagcgtaatt catgagatca gatcggatca gaaaaatcaa
     5461 cagaaaataa acctatttga agtaaaatac ttctaaatac atagtaaact aatgtaatag
     5521 tttttattaa aattacacat atatgtatcg atatatatac atgttttttt ttttttaatt
     5581 ttaaaattat aattgtaatg cactatgcgt aaattgtatt aaatattaat atccaaggca
     5641 tattaataga aatatatata catatataaa acctgcaaaa aaaaaacatg aactaccaag
     5701 ctaaatgttt caataatata tagatctata tgtatgtaaa accgatatga aatctgccta
     5761 cagccagaca aaaaaaacct taaagagaaa gatcagaaga ggaacagaag agtatgagcg
     5821 gggagtgaaa gatgcaagat tcatccaaaa gcgaaaaaca tttttttctt aagtctacgc
     5881 aaaacaaaat gaatgaaaag taacccgaaa ttagttttta gagtataagt ataaaaaatg
     5941 gaaaatatcg cgatagaaag aaacaaaaat ggacgcagat aaggacggca agcagaacaa
     6001 gagtaataaa gcagcggcaa gaactcccta aaaatagcaa agagcattgt aaatatttat
     6061 ataaataatt tatgcctaat taaagtttaa attaatgaag aaaaccagca aaaaaacata
     6121 aaaaaccaga agaaccattt cagatttgta tgcaaatttg tatatgtacg tactttcctt
     6181 taattttagc caatcagttt gtaagattaa attttttttt tgcatctcta taaagaataa
     6241 tgatatgagg ataatgattc cgattttttt ttccgtttaa tgtaaccgtt agaattagcc
     6301 cataatggtc gatatctaaa tgtgtgtgta tatcgatgtg atgtcatcga ttgtcgataa
     6361 cgattggatg cgagtgtgtg ttaatttaat ttgataaaag ataaaggaaa gcaaggaatg
     6421 tgaagataat gaaaggagat gcatagagaa tgagagcgaa tctatctatt atataattat
     6481 gtaaaatgca catctttaaa ttataaatta tacatattta aaaggaaact attattatta
     6541 ttttgattac cattatcatt attttttatt atcacttaac ggtttcggtt tgatccacag
     6601 cgaaaccaac aacaaccaag aaacagaaac aacaaaaaga gaaacaaaat aaacaagcga
     6661 aatgtttgat aaaaaaaacc acgagatgtt ttaatttatt tgtctttaat tcttcgattt
     6721 tattgagttt ttcatagaga aagttgtcac actcaagaac acacacgcaa gcgtataaag
     6781 ctaaacaaat taatacctac taagtgttta aatagtacta aacgaatcaa aatataaata
     6841 cttaaaaaga aaaaaaaacg aaaataaaaa taaaaacagc agcagaaaaa aacgatgaga
     6901 cagttgagca gacgagaaaa acaaagccca ttaaattatt tgttgagtat tttattaaaa
     6961 aataattacg ataataatta aaaagtttaa attgaacgaa ttacgaaaaa caaactgaat
     7021 atacatatat gtgtgcatat atacatatat gtatgtatta tatataaaca aattatttaa
     7081 tagttatgct aaaaaattat ttaaagtgaa cggatatgaa aaatatttgt atgcgtgcga
     7141 gcacggagag aagggcactc agaataggcc atcgacacca cccactccct tccgataccc
     7201 gaacatcacc accagttttc accattttct gacccttaac cccttcaacc agacacacac
     7261 caaacacaag aaaaggcgaa gaaagccgaa aaaaaaacaa tgcaaacacg aaataggcgg
     7321 cgacgacaga gataagcgcg aagaagatac taaatcaata agtaaacata aaaattaatt
     7381 aataaattaa ttccaaaaca aaatgaaaaa agctaaaaaa aaaacatcaa gaaaacaaaa
     7441 gacaaaagca aacaaaaaac atgacgaaac attgaaaata aaaaaatact