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Drosophila melanogaster hormone receptor 4, transcript variant D


LOCUS       NM_001272230            8596 bp    mRNA    linear   INV 26-DEC-2023
            (Hr4), mRNA.
ACCESSION   NM_001272230
VERSION     NM_001272230.1
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 8596)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 8596)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 8596)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 8596)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 8596)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 8596)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 8596)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 8596)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 8596)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 8596)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 8596)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 8596)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 8596)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 8596)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 8596)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 8596)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 8596)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 8596)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..8596
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..8596
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /exception="dicistronic gene"
                     /note="Hormone receptor 4"
                     /map="2B16-2C1"
                     /db_xref="FLYBASE:FBgn0264562"
                     /db_xref="GeneID:31162"
     CDS             1085..5674
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="CG43934 gene product from transcript CG43934-RD;
                     CG43934-PD; Hr4-PD"
                     /codon_start=1
                     /product="hormone receptor 4, isoform D"
                     /protein_id="NP_001259159.1"
                     /db_xref="FLYBASE:FBpp0305532"
                     /db_xref="GeneID:31162"
                     /db_xref="FLYBASE:FBgn0264562"
                     /translation="MPGPALWSGIMTLSRGPYSELDKMSLFQDLKLKRRKIDSRCSSD
                     GESIADTSTSSPDLLAPMSPKLCDSGSAGASLGASLPLPLALPLPMALPLPMSLPLPL
                     TAASSAVTVSLAAVVAAVAETGGAGAGGAGTAVTASGAGPCVSTSSTTAAAATSSTSS
                     LSSSSSSSSSTSSSTSSASPTAGASSTATCPASSSSSSGNGSGGKSGSIKQEHTEIHS
                     SSSAISAAAASTVMSPPPAEATRSSPATPEGGGPAGDGSGATGGGNTSGGSTAGVAIN
                     EHQNNGNGSGGSSRASPDSLEEKPSTTTTTGRPTLTPTNGVLSSASAGTGISTGSSAK
                     LSEAGMSVIRSVKEERLLNVSSKMLVFHQQREQETKAVAAAAAAAAAGHVTVLVTPSR
                     IKSEPPPPASPSSTSSTQRERERERDRERDRERERERDRDREREREQSISSSQQHLSR
                     VSASPPTQLSHGSLGPNIVQTHHLHQQLTQPLTLRKSSPPTEHLLSQSMQHLTQQQAI
                     HLHHLLGQQQQQQQASHPQQQQQQQHSPHSLVRVKKEPNVGQRHLSPHHQQQSPLLQH
                     HQQQQQQQQQQQQHLHQQQQQQQHHQQQPQALALMHPASLALRNSNRDAAILFRVKSE
                     VHQQVAAGLPHLMQSAGGAAAAAAAAVAAQRMVCFSNARINGVKPEVIGGPLGNLRPV
                     GVGGGNGSGSVQCPSPHPSSSSSSSQLSPQTPSQTPPRGTPTVIMGESCGVRTMVWGY
                     EPPPPSAGQSHGQHPQQQQQSPHHQPQQQQQQQQQQSQQQQQQQQQQSLGQQQHCLSS
                     PSAGSLTPSSSSGGGSVSGGGVGGPLTPSSVAPQNNEEAAQLLLSLGQTRIQDMRSRP
                     HPFRTPHALNMERLWAGDYSQLPPGQLQALNLSAQQQQWGSSNSTGLGGVGGGMGGRN
                     LEAPHEPTDEDEQPLVCMICEDKATGLHYGIITCEGCKGFFKRTVQNRRVYTCVADGT
                     CEITKAQRNRCQYCRFKKCIEQGMVLQAVREDRMPGGRNSGAVYNLYKVKYKKHKKTN
                     QKQQQQAAQQQQQQAAAQQQHQQQQQHQQHQQHQQQQLHSPLHHHHHQGHQSHHAQQQ
                     HHPQLSPHHLLSPQQQQLAAAVAAAAQHQQQQQQQQQQQQQAKLMGGVVDMKPMFLGP
                     ALKPELLQAPPMHSPAQQQQQQQQQQQQQQASPHLSLSSPHQQQQQQQGQHQNHHQQQ
                     GGGGGGAGGGAQLPPHLVNGTILKTALTNPSEIVHLRHRLDSAVSSSKDRQISYEHAL
                     GMIQTLIDCDAMEDIATLPHFSEFLEDKSEISEKLCNIGDSIVHKLVSWTKKLPFYLE
                     IPVEIHTKLLTDKWHEILILTTAAYQALHGKRRGEGGGSRHGSPASTPLSTPTGTPLS
                     TPIPSPAQPLHKDDPEFVSEVNSHLSTLQTCLTTLMGQPIAMEQLKLDVGHMVDKMTQ
                     ITIMFRRIKLKMEEYVCLKVYILLNKAEVELESIQERYVQVLRSYLQNSSPQNPQARL
                     SELLSHIPEIQAAASLLLESKMFYVPFVLNSASIR"
     misc_feature    3851..4120
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="DNA-binding domain of ecdysone-induced DHR4 orphan
                     nuclear receptor is composed of two C4-type zinc fingers;
                     Region: NR_DBD_DHR4_like; cd07168"
                     /db_xref="CDD:143542"
     misc_feature    order(3875..3877,3884..3886,3926..3928,3935..3937,
                     3983..3985,4001..4003,4031..4033,4040..4042)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="zinc binding site [ion binding]; other site"
                     /db_xref="CDD:143542"
     misc_feature    order(3905..3913,3929..3934,3938..3940,3944..3946,
                     3953..3955,3962..3964,4022..4027,4034..4036,4043..4045,
                     4076..4078,4085..4087)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative DNA binding site [nucleotide binding];
                     other site"
                     /db_xref="CDD:143542"
     misc_feature    4880..5659
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="The ligand binding domain of orphan nuclear
                     receptor Ecdysone-induced receptor DHR4; Region:
                     NR_LBD_DHR4_like; cd06953"
                     /db_xref="CDD:132751"
     misc_feature    order(4970..4975,4985..4987,5087..5089,5099..5101,
                     5108..5110,5306..5308,5375..5377)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative ligand binding site [chemical binding];
                     other site"
                     /db_xref="CDD:132751"
     misc_feature    order(5000..5002,5009..5011,5021..5023,5036..5038,
                     5051..5056,5060..5065,5072..5077)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative coactivator recognition site [active]"
                     /db_xref="CDD:132751"
ORIGIN      
        1 gtgtatccca cgagctagtc gcgcgcggaa agcaaacgct tcgaaaatca aacgatattt
       61 ccgttttgac gtttaacccc cacccaccca aataaagcaa atctaactaa agagtcctgc
      121 aaaaaaatca acagctgctt gatcttgaag cctccatatc ttgaaatcct gcatcagcca
      181 acagccccaa aacaaaacaa aacaaaacga atgcaaatac actaatgtga acaacaacta
      241 aattgctagt gaaatgtttt aagccaaatg ctgtgagtgt gtaacaaaaa aaaaaaaaaa
      301 aaaaaaaaaa aaagaagagt gattcaaagc aaccgcaagc aatcaacttg taacggtagt
      361 cggatactcg cttgcccgcg gagcatggat atcgccccag ttccgccacc accattatcg
      421 atatccgtat cgctttcggc tccaggatca acggtattcg caaacgctgg ccactagaga
      481 agcgtcagcc gaatgcctca aagatgacac cgctgcaacg ggaccaattg aaccaccaac
      541 aatttgtgca ccactatttg caaagtcaac ctgtcaataa caaaattgac agctttgata
      601 ttaaacatat acgtaaaacc caaaacccga ccacaaaccc aaagtttcga actgaagacc
      661 gtttcaagac aagcagcaag agcaagatta ccaacccgga gagcggaaat acccaactcc
      721 attttcattt gagtgaaacg ccagacttat ccgcctcaac ttcgctgttg ccgtcagcat
      781 ttgttgcatc ctccgccgcc ttggtgaatg tgacatgcca ggcggaggaa ggacaaagtt
      841 cagcaggatc acactacaca gttcatagag ggaatgacgc aggcatttct tttgcagcgt
      901 tggactttcg ttttataagt gagttctaga cttgagtgct tgcatactat gtggcagcca
      961 ctggcgcccg cactggccca ctgaaaagga aagagtgcga aaccaggccg aaataaaagt
     1021 ttgcgagcaa acaacagcag cccccaagac gtcaggcaaa cgacaacaac aacaacaacc
     1081 agctatgcct ggcccggctt tgtggtccgg aattatgaca ctgagccgtg gcccgtacag
     1141 cgagctcgat aaaatgagcc tttttcaaga cctcaaactc aaacggcgca aaatcgattc
     1201 gcgatgcagc agtgacggcg agtccatagc ggacacgtcc acctcgtcgc cggacctgct
     1261 ggcgcccatg tcgccgaagc tctgcgacag cggctcggcg ggggcgtcgc tgggggcatc
     1321 gctgcccctg ccgctggccc tgcccctgcc aatggccctg ccactgccca tgtcgctgcc
     1381 cctgcccctc acggcggcat cttcggcggt caccgtttcg ctggcagcgg tcgtggccgc
     1441 ggtggccgag acgggtggcg cgggcgcggg aggagctggg acagcagtaa cagcgtcggg
     1501 agcaggacca tgcgtctcca cgtcgtctac gacggcagcg gcagccacat cctcgacctc
     1561 ctcgctctcg tcctcctcct cttcgtcatc ctccacgtcc tccagcactt cctccgcctc
     1621 gccgacagct ggagcctcct ccacggccac ctgccccgcc agcagcagca gcagcagtgg
     1681 aaacggaagt gggggcaaaa gtggtagcat caagcaggag cacacggaga tacactcgtc
     1741 gagcagtgcg atttcggcgg ccgccgcctc aacggtgatg tcaccgccgc ccgctgaggc
     1801 gacgagatcc agtccagcca cgcccgaggg aggcggacca gctggcgacg gaagtggagc
     1861 aacgggaggc ggaaacacga gcggcggatc aacggctgga gtggccatta atgaacacca
     1921 aaacaatggc aatggcagcg gcgggagcag tcgagcctct cccgattcgc tggaagagaa
     1981 gccctctacc acaacgacca caggtcgtcc aacgctcacg cccacgaatg gggtgctgtc
     2041 ctccgcctcg gcgggcacgg ggatttccac aggaagcagc gccaagctga gcgaggctgg
     2101 tatgagtgtg atacggtccg tgaaggagga gcgcttgctc aacgtatcca gcaagatgct
     2161 ggtgttccat cagcagcggg agcaagagac caaagcagtg gcggctgcag cagcagcagc
     2221 agcggcgggc catgtgacgg ttctagtgac gccatcgcgc atcaaatcgg agccaccgcc
     2281 gccggcttca ccctcctcta catccagcac acaaagggaa agggaacggg aacgcgatcg
     2341 agagagggat cgcgaaaggg aacgcgagcg ggaccgggac cgggaacggg aacgggaaca
     2401 gtccatcagc tcctcgcagc agcacctaag tcgggtctcc gccagtccac ccactcagct
     2461 gtcccacggc agcctgggac ccaacattgt gcagacgcac catcttcacc agcaactcac
     2521 acagccgctg acgctgcgca agagcagccc gcccacagag cacctgctca gtcagtccat
     2581 gcaacatctc acacagcagc aggcgatcca cctgcatcac ctacttggcc agcagcagca
     2641 gcagcagcag gcgtcgcatc cccagcagca acagcagcag caacactcgc cccactccct
     2701 ggtgcgggtg aaaaaggaac cgaatgttgg tcagcggcac ttatcgccgc atcaccaaca
     2761 acagtcgcca ctcctgcagc accaccaaca gcagcagcag cagcaacaac aacagcaaca
     2821 gcatctgcat cagcaacagc aacagcagca gcatcaccag cagcagcccc aggcactggc
     2881 cctgatgcat ccggcttccc tggcgctaag gaacagcaat cgggatgcgg ccattctgtt
     2941 tcgggtgaag agcgaagtgc accagcaggt ggccgccggg ctgccgcatc tgatgcagtc
     3001 cgctggtggg gcagcggccg ccgccgcagc agctgtggcc gctcagcgaa tggtatgctt
     3061 cagcaatgcc aggatcaatg gcgttaagcc ggaggtgatt ggaggaccgc tgggcaacct
     3121 gcggcccgtg ggcgtcggtg gcggaaacgg aagtggctcc gtgcagtgcc cctcgccgca
     3181 tccatcctcc tcgtcgtcat cctcgcagct gtcgccgcag acgccctccc agacgccgcc
     3241 ccgaggcacg cccaccgtca taatgggcga gagctgcggg gtgcgcacca tggtctgggg
     3301 ctacgagcct ccgccaccct cggcgggcca gtcccacggc cagcacccgc aacagcaaca
     3361 gcagtcgccc caccaccagc cgcaacaaca acagcagcag caacaacagc agtcgcagca
     3421 gcaacagcaa cagcagcagc aacagtcgct gggccagcag cagcactgcc tctcctcgcc
     3481 gtcggcggga tcgctgacgc cctcctcttc gtccggcggt ggttcggtat ctggcggcgg
     3541 agtgggcgga ccactcacac cctcctcggt ggcgccgcag aataacgagg aggccgccca
     3601 actcctgctc tccctgggac agacacgcat ccaggacatg agatcacggc cacacccctt
     3661 ccgcacaccg cacgccctta atatggagcg gctgtgggcg ggagactact cgcaattgcc
     3721 gcccggccag ctgcaggctc tgaatctcag tgcccaacag cagcagtggg gcagcagcaa
     3781 ctccacgggt cttggtggcg taggcggcgg catgggcgga cgcaacctgg aggcgccgca
     3841 cgagccgacc gacgaggacg aacagccgct cgtttgcatg atctgcgagg acaaggccac
     3901 cggcctgcac tacggcatca tcacctgcga ggggtgcaag ggcttcttca agcggacggt
     3961 gcagaaccga cgagtctaca cctgcgtggc ggacggcacc tgcgagataa ccaaagcaca
     4021 gcgcaaccgt tgtcagtatt gtcgatttaa gaagtgcatc gagcagggca tggtgctgca
     4081 agccgttcgc gaggatcgca tgccgggcgg tcgcaacagt ggcgccgtct acaatttgta
     4141 caaggtgaag tacaagaagc acaagaagac caatcagaag cagcagcagc aggccgccca
     4201 gcagcagcag cagcaggcgg cggcgcagca gcagcaccag caacagcagc agcatcaaca
     4261 gcaccagcaa catcagcaac agcagttgca ctcgccgctc caccatcacc accaccaggg
     4321 ccaccagtcg caccacgcgc agcagcagca ccacccacag ctgtcgccgc accacctgct
     4381 gtcgccgcag cagcagcaac ttgccgccgc ggtggcagca gctgcgcagc accaacagca
     4441 acagcaacaa cagcagcaac agcagcagca ggccaagctg atgggcggcg tggtggacat
     4501 gaagcccatg ttcctcggcc ccgctttgaa gccggagttg ctgcaagcac cccccatgca
     4561 cagtccggcc cagcaacaac aacagcagca gcagcagcag cagcaacagc aggcctcgcc
     4621 gcatctctcg cttagctcac cgcaccagca gcagcagcag cagcagggac agcaccaaaa
     4681 ccaccaccag caacaaggtg ggggtggcgg aggagctggt ggaggagctc aactgccgcc
     4741 gcacctggtg aacggaacga tactgaagac ggccctaacc aatcccagcg agattgtaca
     4801 tctgcgccac cgcctcgact cggcggtcag ttcgtccaag gaccgacaga tctcgtacga
     4861 gcacgcctta ggcatgatcc agacactgat cgactgcgac gcgatggagg acatagccac
     4921 actgccgcac ttcagcgagt tccttgagga caagtcggag attagcgaga aactgtgcaa
     4981 catcggcgat tccatagtcc acaagctggt gtcgtggaca aaaaagttgc ccttctacct
     5041 ggagatcccg gtggagatac ataccaaact actgacggac aagtggcacg agatccttat
     5101 cctgaccacg gccgcctacc aggcgttgca tggcaagcgg cgtggcgagg gaggaggcag
     5161 caggcatggt tcgccggcgt caacgccact gagcacgccc actggtacgc cgttgagcac
     5221 accgataccc tcgcccgccc agccactgca caaggacgac ccggagtttg tcagcgaggt
     5281 gaactcgcac ctgagcacac tgcaaacctg cttgaccacg ctaatgggcc agccgatagc
     5341 gatggagcag ctgaagctgg acgtcgggca catggtggac aagatgaccc agatcaccat
     5401 catgttccgg cgaatcaagc tcaagatgga ggagtacgtc tgcctgaagg tttacatact
     5461 gctaaacaaa gcagaagtgg aactggagag catccaggag cggtacgtcc aggtgctgcg
     5521 ctcctacctg caaaactcct cgccgcagaa tccgcaggcg aggctcagtg aactgctctc
     5581 ccacatacca gagatccagg ctgcggctag cctgctgctc gagagcaaga tgttctatgt
     5641 gcccttcgtg ctcaactcgg cgagcataag gtagcagatg cttgagcacg gactgctgcc
     5701 caccgccaac catcagtcgg cgccaccacc tggtcctgat acgatcccgg agcggcagca
     5761 gcaacagcaa caggagcggc gtcgaagcag gcagcagcag caccagttgc tgcgcaacct
     5821 gcccaagatc aagatcgaga tcggttccca gcgcgatccg gaggaagagg agtcggagca
     5881 agagctagag caggagccgg agcaagagca ggatgtgcag tcccgaccct cagccgcatc
     5941 cacgtccatt ttgaagactt cactgctgag cggagcggcg gcgacgttgg ccaccttgac
     6001 ggcggcagcc gagcaaatag cccgctcctc ggccagctgc gacactgcca gggattattc
     6061 ccaaagcagc aacagcagca gcaatagcaa cgctgccaca ggagcaacag gaactatggc
     6121 cagcggcagg acaggcagca gcaacagttt agaagagagc aaaagacttg agcatcaaca
     6181 acaacaacaa caacagcagc aacaaccaca acagcatcag cagcaacaat atacatcaat
     6241 actacacacg gccttaatgt cacagcgttc catgccacca gcgacatcag ctacatcagc
     6301 agcagcggcg gtcacatcga cgccgacaac ggcagcaaca acatcgccta cgcctgcgca
     6361 tgtgctgaaa acagcggtga ccttagccag tctaagtgag atagccacag cacgtcagca
     6421 gcagcaggag cagcagcagc aacagcagca gcagcagcag cagcaacaat tgctggcgat
     6481 gcaacagcaa cagcaacagc taacgaaact gcccagaagg ataatcaact ttggcagcaa
     6541 tcacactgcc aacactgcca ccaaagcatt gggagcggga tcagaagcag gagcaggagc
     6601 aggagtgggc atggccacag caacggcaac agcaacagtt ggacgcaatc gccaactggg
     6661 cctgagtcag ggtcaagtgg tgaatggtaa gcagaaagaa ttgtgagcct tgtgtttgcg
     6721 ataacgtttg tgtacccctc gttaacgctg ccagtcagga cagcacgcct gctcaatggc
     6781 aactgttgcg gccgattgca ggcgacagca ggaacagggg ccagcagggg caccaacact
     6841 gccggcagca acacggtcaa catgcagacc cagaccgacg atgagttgct gccgctgtta
     6901 gccatgctcc cacaagacgc tcccacaaca acaggagcag cagcaggagc agcagtatca
     6961 ttatcattat tagccacagc cacagccaca accacatctt catcatcatc atcatcatca
     7021 tcattatcat catcatcgac gatttcatta gcaacacgtg cagcagcagt aatgcagcag
     7081 cagcaacaat ggactgccgt gtcgacgacg ccagcaacag gagtgatgat gacgacgacg
     7141 acgacacaat cacaatcaca agcaacatta gcaccacaac aacagcaact atcactgcca
     7201 caacaaaagc aacaatttaa tacgagaata aacagaaaaa caacaattac aacagccacc
     7261 acaacagcac caacagcaac aacagcaaca acaacaacaa cacgaaacag caacatcaac
     7321 ttcaacagca acaataacta tatgaataac aataatataa acgccgccac tgccagagcc
     7381 tctgccttcg cctccgcctc tgccatggcc acggccacgc ccacaggcac acccactgcc
     7441 acgggcccgc gccagagacc ggccccgcaa ccgcccccgc ccccgcgcac gcccacagcc
     7501 acctcagctt ctgttactgt agttgtgttc aaaacaatgt tacctgatga ctgaattatt
     7561 ttagaatcgt acacatagag catgttttat tatgattacc actatgatta ccacttacga
     7621 tttacgatta cgattatgat tacgattatg agtatgtctg gatatcatta cctattaact
     7681 attcattagt tatggattat cgattattac tatttttact attactattg ctacatgcat
     7741 aatgtagagc ctaattgatc tatcgtgtat tttagtttgt ctaattgttg gatccctgta
     7801 ctatatccca cacacataca cacactctca cccatatttg atcctcaccc ccaaacaatc
     7861 cataagcaca cactcataca cacacacaca cacacacaca cattgccgtt actaaggaag
     7921 ggtatgcgca tatgcgaagt tgaaacattt taatcatgta tttttttttt ttgtgttttt
     7981 gctgaacgga gatgaaagtt aaggaggtcg agcaacagca attagaagtg ttaattgagt
     8041 aggattatga gtacgtagcg agtatgtaac gagtatgtaa tgactaagta aagtttacgt
     8101 agacacaatg ccacacacag ctatagcaag catttaggat gcgtgcatat gtcagttatc
     8161 tatttttttt ttttaaacta aaatctatat aaagtaacga gagaggaagg aagagcgaag
     8221 cagggcggga gattcaacca cataaagtat aaatagtaga gcttgagcca agtagctagt
     8281 ttagtgtaca tccgttttgg ggcagggaag tagagcaatt tgttgacgag tcggggtggt
     8341 ggggtgagcg gtaacgaaga tcggtaacgg aagtggagac aaagcggaac cggaaacggg
     8401 agctggactg ccagatcgga atcacacatc aaccattatg agaacttcgc tcgcaactct
     8461 tgcagaatta tactgttaga tcctgttagt tatccaaccc aaaactccca acccacacac
     8521 actataccta agttaagaac gcataatcac atactacaca taaattacgc atagtacata
     8581 caaaaaagat gaagag