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Drosophila melanogaster hormone receptor 4, transcript variant E


LOCUS       NM_001272229            9374 bp    mRNA    linear   INV 26-DEC-2023
            (Hr4), mRNA.
ACCESSION   NM_001272229
VERSION     NM_001272229.1
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 9374)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 9374)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 9374)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 9374)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 9374)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 9374)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 9374)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 9374)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 9374)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 9374)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 9374)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 9374)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 9374)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 9374)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 9374)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 9374)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 9374)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 9374)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..9374
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..9374
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /exception="dicistronic gene"
                     /note="Hormone receptor 4"
                     /map="2B16-2C1"
                     /db_xref="FLYBASE:FBgn0264562"
                     /db_xref="GeneID:31162"
     CDS             924..6452
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="CG43934 gene product from transcript CG43934-RE;
                     CG43934-PE; Hr4-PE"
                     /codon_start=1
                     /product="hormone receptor 4, isoform E"
                     /protein_id="NP_001259158.1"
                     /db_xref="FLYBASE:FBpp0305531"
                     /db_xref="GeneID:31162"
                     /db_xref="FLYBASE:FBgn0264562"
                     /translation="MNAQQPLILEPGSKASSVVAAAAELEAATTTAALATSSATEASA
                     ADSYPAVQQNGGHEEQRLNGNHGCRREQPDDRTDQPEQLDQQDEPEPVKDVANKLTII
                     KTPLNKTLLKKCNSNGSLQMPHANNGSSININNRNNTNNTNNNNSNNNNNAASNDVCD
                     SGLQSGNSNEQEPFANNDRHHHIYHHHHHHYHHHHGGDKSEASGDVASSSSADPNSQV
                     KDEPTAQDSQDNQAAGAAEAGGATAVAATCSPSKCNKCNSHNNNSNCNNNNNTISNNN
                     NNTSSNTNTNSNSNNNANNNNNNNNNNHYNNYNNYYKKKSRMPPICRIMTLSRGPYSE
                     LDKMSLFQDLKLKRRKIDSRCSSDGESIADTSTSSPDLLAPMSPKLCDSGSAGASLGA
                     SLPLPLALPLPMALPLPMSLPLPLTAASSAVTVSLAAVVAAVAETGGAGAGGAGTAVT
                     ASGAGPCVSTSSTTAAAATSSTSSLSSSSSSSSSTSSSTSSASPTAGASSTATCPASS
                     SSSSGNGSGGKSGSIKQEHTEIHSSSSAISAAAASTVMSPPPAEATRSSPATPEGGGP
                     AGDGSGATGGGNTSGGSTAGVAINEHQNNGNGSGGSSRASPDSLEEKPSTTTTTGRPT
                     LTPTNGVLSSASAGTGISTGSSAKLSEAGMSVIRSVKEERLLNVSSKMLVFHQQREQE
                     TKAVAAAAAAAAAGHVTVLVTPSRIKSEPPPPASPSSTSSTQRERERERDRERDRERE
                     RERDRDREREREQSISSSQQHLSRVSASPPTQLSHGSLGPNIVQTHHLHQQLTQPLTL
                     RKSSPPTEHLLSQSMQHLTQQQAIHLHHLLGQQQQQQQASHPQQQQQQQHSPHSLVRV
                     KKEPNVGQRHLSPHHQQQSPLLQHHQQQQQQQQQQQQHLHQQQQQQQHHQQQPQALAL
                     MHPASLALRNSNRDAAILFRVKSEVHQQVAAGLPHLMQSAGGAAAAAAAAVAAQRMVC
                     FSNARINGVKPEVIGGPLGNLRPVGVGGGNGSGSVQCPSPHPSSSSSSSQLSPQTPSQ
                     TPPRGTPTVIMGESCGVRTMVWGYEPPPPSAGQSHGQHPQQQQQSPHHQPQQQQQQQQ
                     QQSQQQQQQQQQQSLGQQQHCLSSPSAGSLTPSSSSGGGSVSGGGVGGPLTPSSVAPQ
                     NNEEAAQLLLSLGQTRIQDMRSRPHPFRTPHALNMERLWAGDYSQLPPGQLQALNLSA
                     QQQQWGSSNSTGLGGVGGGMGGRNLEAPHEPTDEDEQPLVCMICEDKATGLHYGIITC
                     EGCKGFFKRTVQNRRVYTCVADGTCEITKAQRNRCQYCRFKKCIEQGMVLQAVREDRM
                     PGGRNSGAVYNLYKVKYKKHKKTNQKQQQQAAQQQQQQAAAQQQHQQQQQHQQHQQHQ
                     QQQLHSPLHHHHHQGHQSHHAQQQHHPQLSPHHLLSPQQQQLAAAVAAAAQHQQQQQQ
                     QQQQQQQAKLMGGVVDMKPMFLGPALKPELLQAPPMHSPAQQQQQQQQQQQQQQASPH
                     LSLSSPHQQQQQQQGQHQNHHQQQGGGGGGAGGGAQLPPHLVNGTILKTALTNPSEIV
                     HLRHRLDSAVSSSKDRQISYEHALGMIQTLIDCDAMEDIATLPHFSEFLEDKSEISEK
                     LCNIGDSIVHKLVSWTKKLPFYLEIPVEIHTKLLTDKWHEILILTTAAYQALHGKRRG
                     EGGGSRHGSPASTPLSTPTGTPLSTPIPSPAQPLHKDDPEFVSEVNSHLSTLQTCLTT
                     LMGQPIAMEQLKLDVGHMVDKMTQITIMFRRIKLKMEEYVCLKVYILLNKEVELESIQ
                     ERYVQVLRSYLQNSSPQNPQARLSELLSHIPEIQAAASLLLESKMFYVPFVLNSASIR
                     "
     misc_feature    4632..4901
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="DNA-binding domain of ecdysone-induced DHR4 orphan
                     nuclear receptor is composed of two C4-type zinc fingers;
                     Region: NR_DBD_DHR4_like; cd07168"
                     /db_xref="CDD:143542"
     misc_feature    order(4656..4658,4665..4667,4707..4709,4716..4718,
                     4764..4766,4782..4784,4812..4814,4821..4823)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="zinc binding site [ion binding]; other site"
                     /db_xref="CDD:143542"
     misc_feature    order(4686..4694,4710..4715,4719..4721,4725..4727,
                     4734..4736,4743..4745,4803..4808,4815..4817,4824..4826,
                     4857..4859,4866..4868)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative DNA binding site [nucleotide binding];
                     other site"
                     /db_xref="CDD:143542"
     misc_feature    5661..6437
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="The ligand binding domain of orphan nuclear
                     receptor Ecdysone-induced receptor DHR4; Region:
                     NR_LBD_DHR4_like; cd06953"
                     /db_xref="CDD:132751"
     misc_feature    order(5751..5756,5766..5768,5868..5870,5880..5882,
                     5889..5891,6087..6089,6156..6158)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative ligand binding site [chemical binding];
                     other site"
                     /db_xref="CDD:132751"
     misc_feature    order(5781..5783,5790..5792,5802..5804,5817..5819,
                     5832..5837,5841..5846,5853..5858)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative coactivator recognition site [active]"
                     /db_xref="CDD:132751"
ORIGIN      
        1 tttcacttta tttcgttcgc atgcacgcac gttggccgcg gagcaacatc tccagcaaac
       61 agaaatcgaa acctaaacag aaacagaaac agaagcaaaa aataaaacat aaaaaaaaaa
      121 caaaaaaaaa taaaaaacaa aacgaaacta caaatacaaa taacaaaaaa caaataacac
      181 ggcagcagcc gtcgctgaga cgcgaaactt ggctcgaact ttagtttagt ttttctcagt
      241 ttcagtggcc agtggctaaa gtgcggcgat atcgggcgaa acggttaacg gctcacggct
      301 catggctaac ggttgacagc cagcggttca tcacgcgcca aatccttttt tttcaagttt
      361 gtcaatgttt ttagtggtac taaagaaaac acatcattat atgtaaacta taggcaagaa
      421 tctaaagcca agagtggtat gtgcccttag ttttgaccca agacctacaa gtccctagaa
      481 gacccctgac agccagaacc acccacccac acaccacgca taaacacacg cacacacaca
      541 aacacactca ttcgccaccc acccacacag tctcatatct cagtctcagt ctcagtttca
      601 atctcagttc caatctcagt tccaatccaa tctcaatctc atttcgagga tcaaaaacgg
      661 agcagcaaca tcaatcgatt cattatgttt acatattttt ttcgagtgat ttaagttttg
      721 gcaatggacg cagacttctt ctccatctaa tcgaaccgag tgacttcgca tcatgctgta
      781 gaagaagaag ttgactaaca aatggttgtt gtgatcgaaa gtgtcttttg agatgtgttt
      841 aaaactcggt gtttgcccag ccaaaatggc ccagcatcgt tcgcttgtcc gtctctacct
      901 gcccacctag aagagctgcc cagatgaatg cccagcagcc gctgatcctg gagcctggat
      961 ccaaggccag cagtgttgtt gcggcagctg ccgagctaga ggcagcaaca accacggcag
     1021 cgctggccac gtcctccgcc acggaggctt ctgcagcgga cagctatcct gccgtacagc
     1081 agaacggcgg ccacgaggag cagcggctca atggcaacca cggatgtcgc cgcgagcagc
     1141 cggatgatcg gacggaccag ccggagcagt tggatcagca ggatgagccc gagccagtca
     1201 aggacgtggc caacaagctg accattatca agacgccgtt gaacaagact ctgctgaaga
     1261 agtgcaacag caacgggagc ctgcagatgc cgcacgccaa caatggcagc agcatcaaca
     1321 tcaataatcg caacaacaca aacaacacca acaacaacaa tagcaacaat aacaacaatg
     1381 cggccagcaa cgacgtctgc gattccggtt tgcaaagcgg aaatagcaat gaacaagaac
     1441 catttgccaa caacgatcgc catcaccaca tctaccacca ccaccaccat cactatcatc
     1501 atcaccatgg cggcgataag agcgaggcga gcggcgacgt tgcatcctcg tcctccgcgg
     1561 accccaactc gcaggtcaag gacgagccca cagcgcagga tagccaggac aatcaagcgg
     1621 ccggagctgc agaggctgga ggagctactg cggttgcagc cacgtgcagc cccagcaagt
     1681 gcaacaagtg caactcgcac aataacaaca gcaactgcaa caataacaat aataccatca
     1741 gcaacaacaa caataataca agcagtaata ccaatacgaa cagcaacagc aacaacaatg
     1801 ccaacaacaa taataataat aacaataata accattacaa caactacaat aactattata
     1861 agaaaaagtc tcgtatgccg cccatttgcc gaattatgac actgagccgt ggcccgtaca
     1921 gcgagctcga taaaatgagc ctttttcaag acctcaaact caaacggcgc aaaatcgatt
     1981 cgcgatgcag cagtgacggc gagtccatag cggacacgtc cacctcgtcg ccggacctgc
     2041 tggcgcccat gtcgccgaag ctctgcgaca gcggctcggc gggggcgtcg ctgggggcat
     2101 cgctgcccct gccgctggcc ctgcccctgc caatggccct gccactgccc atgtcgctgc
     2161 ccctgcccct cacggcggca tcttcggcgg tcaccgtttc gctggcagcg gtcgtggccg
     2221 cggtggccga gacgggtggc gcgggcgcgg gaggagctgg gacagcagta acagcgtcgg
     2281 gagcaggacc atgcgtctcc acgtcgtcta cgacggcagc ggcagccaca tcctcgacct
     2341 cctcgctctc gtcctcctcc tcttcgtcat cctccacgtc ctccagcact tcctccgcct
     2401 cgccgacagc tggagcctcc tccacggcca cctgccccgc cagcagcagc agcagcagtg
     2461 gaaacggaag tgggggcaaa agtggtagca tcaagcagga gcacacggag atacactcgt
     2521 cgagcagtgc gatttcggcg gccgccgcct caacggtgat gtcaccgccg cccgctgagg
     2581 cgacgagatc cagtccagcc acgcccgagg gaggcggacc agctggcgac ggaagtggag
     2641 caacgggagg cggaaacacg agcggcggat caacggctgg agtggccatt aatgaacacc
     2701 aaaacaatgg caatggcagc ggcgggagca gtcgagcctc tcccgattcg ctggaagaga
     2761 agccctctac cacaacgacc acaggtcgtc caacgctcac gcccacgaat ggggtgctgt
     2821 cctccgcctc ggcgggcacg gggatttcca caggaagcag cgccaagctg agcgaggctg
     2881 gtatgagtgt gatacggtcc gtgaaggagg agcgcttgct caacgtatcc agcaagatgc
     2941 tggtgttcca tcagcagcgg gagcaagaga ccaaagcagt ggcggctgca gcagcagcag
     3001 cagcggcggg ccatgtgacg gttctagtga cgccatcgcg catcaaatcg gagccaccgc
     3061 cgccggcttc accctcctct acatccagca cacaaaggga aagggaacgg gaacgcgatc
     3121 gagagaggga tcgcgaaagg gaacgcgagc gggaccggga ccgggaacgg gaacgggaac
     3181 agtccatcag ctcctcgcag cagcacctaa gtcgggtctc cgccagtcca cccactcagc
     3241 tgtcccacgg cagcctggga cccaacattg tgcagacgca ccatcttcac cagcaactca
     3301 cacagccgct gacgctgcgc aagagcagcc cgcccacaga gcacctgctc agtcagtcca
     3361 tgcaacatct cacacagcag caggcgatcc acctgcatca cctacttggc cagcagcagc
     3421 agcagcagca ggcgtcgcat ccccagcagc aacagcagca gcaacactcg ccccactccc
     3481 tggtgcgggt gaaaaaggaa ccgaatgttg gtcagcggca cttatcgccg catcaccaac
     3541 aacagtcgcc actcctgcag caccaccaac agcagcagca gcagcaacaa caacagcaac
     3601 agcatctgca tcagcaacag caacagcagc agcatcacca gcagcagccc caggcactgg
     3661 ccctgatgca tccggcttcc ctggcgctaa ggaacagcaa tcgggatgcg gccattctgt
     3721 ttcgggtgaa gagcgaagtg caccagcagg tggccgccgg gctgccgcat ctgatgcagt
     3781 ccgctggtgg ggcagcggcc gccgccgcag cagctgtggc cgctcagcga atggtatgct
     3841 tcagcaatgc caggatcaat ggcgttaagc cggaggtgat tggaggaccg ctgggcaacc
     3901 tgcggcccgt gggcgtcggt ggcggaaacg gaagtggctc cgtgcagtgc ccctcgccgc
     3961 atccatcctc ctcgtcgtca tcctcgcagc tgtcgccgca gacgccctcc cagacgccgc
     4021 cccgaggcac gcccaccgtc ataatgggcg agagctgcgg ggtgcgcacc atggtctggg
     4081 gctacgagcc tccgccaccc tcggcgggcc agtcccacgg ccagcacccg caacagcaac
     4141 agcagtcgcc ccaccaccag ccgcaacaac aacagcagca gcaacaacag cagtcgcagc
     4201 agcaacagca acagcagcag caacagtcgc tgggccagca gcagcactgc ctctcctcgc
     4261 cgtcggcggg atcgctgacg ccctcctctt cgtccggcgg tggttcggta tctggcggcg
     4321 gagtgggcgg accactcaca ccctcctcgg tggcgccgca gaataacgag gaggccgccc
     4381 aactcctgct ctccctggga cagacacgca tccaggacat gagatcacgg ccacacccct
     4441 tccgcacacc gcacgccctt aatatggagc ggctgtgggc gggagactac tcgcaattgc
     4501 cgcccggcca gctgcaggct ctgaatctca gtgcccaaca gcagcagtgg ggcagcagca
     4561 actccacggg tcttggtggc gtaggcggcg gcatgggcgg acgcaacctg gaggcgccgc
     4621 acgagccgac cgacgaggac gaacagccgc tcgtttgcat gatctgcgag gacaaggcca
     4681 ccggcctgca ctacggcatc atcacctgcg aggggtgcaa gggcttcttc aagcggacgg
     4741 tgcagaaccg acgagtctac acctgcgtgg cggacggcac ctgcgagata accaaagcac
     4801 agcgcaaccg ttgtcagtat tgtcgattta agaagtgcat cgagcagggc atggtgctgc
     4861 aagccgttcg cgaggatcgc atgccgggcg gtcgcaacag tggcgccgtc tacaatttgt
     4921 acaaggtgaa gtacaagaag cacaagaaga ccaatcagaa gcagcagcag caggccgccc
     4981 agcagcagca gcagcaggcg gcggcgcagc agcagcacca gcaacagcag cagcatcaac
     5041 agcaccagca acatcagcaa cagcagttgc actcgccgct ccaccatcac caccaccagg
     5101 gccaccagtc gcaccacgcg cagcagcagc accacccaca gctgtcgccg caccacctgc
     5161 tgtcgccgca gcagcagcaa cttgccgccg cggtggcagc agctgcgcag caccaacagc
     5221 aacagcaaca acagcagcaa cagcagcagc aggccaagct gatgggcggc gtggtggaca
     5281 tgaagcccat gttcctcggc cccgctttga agccggagtt gctgcaagca ccccccatgc
     5341 acagtccggc ccagcaacaa caacagcagc agcagcagca gcagcaacag caggcctcgc
     5401 cgcatctctc gcttagctca ccgcaccagc agcagcagca gcagcaggga cagcaccaaa
     5461 accaccacca gcaacaaggt gggggtggcg gaggagctgg tggaggagct caactgccgc
     5521 cgcacctggt gaacggaacg atactgaaga cggccctaac caatcccagc gagattgtac
     5581 atctgcgcca ccgcctcgac tcggcggtca gttcgtccaa ggaccgacag atctcgtacg
     5641 agcacgcctt aggcatgatc cagacactga tcgactgcga cgcgatggag gacatagcca
     5701 cactgccgca cttcagcgag ttccttgagg acaagtcgga gattagcgag aaactgtgca
     5761 acatcggcga ttccatagtc cacaagctgg tgtcgtggac aaaaaagttg cccttctacc
     5821 tggagatccc ggtggagata cataccaaac tactgacgga caagtggcac gagatcctta
     5881 tcctgaccac ggccgcctac caggcgttgc atggcaagcg gcgtggcgag ggaggaggca
     5941 gcaggcatgg ttcgccggcg tcaacgccac tgagcacgcc cactggtacg ccgttgagca
     6001 caccgatacc ctcgcccgcc cagccactgc acaaggacga cccggagttt gtcagcgagg
     6061 tgaactcgca cctgagcaca ctgcaaacct gcttgaccac gctaatgggc cagccgatag
     6121 cgatggagca gctgaagctg gacgtcgggc acatggtgga caagatgacc cagatcacca
     6181 tcatgttccg gcgaatcaag ctcaagatgg aggagtacgt ctgcctgaag gtttacatac
     6241 tgctaaacaa agaagtggaa ctggagagca tccaggagcg gtacgtccag gtgctgcgct
     6301 cctacctgca aaactcctcg ccgcagaatc cgcaggcgag gctcagtgaa ctgctctccc
     6361 acataccaga gatccaggct gcggctagcc tgctgctcga gagcaagatg ttctatgtgc
     6421 ccttcgtgct caactcggcg agcataaggt agcagatgct tgagcacgga ctgctgccca
     6481 ccgccaacca tcagtcggcg ccaccacctg gtcctgatac gatcccggag cggcagcagc
     6541 aacagcaaca ggagcggcgt cgaagcaggc agcagcagca ccagttgctg cgcaacctgc
     6601 ccaagatcaa gatcgagatc ggttcccagc gcgatccgga ggaagaggag tcggagcaag
     6661 agctagagca ggagccggag caagagcagg atgtgcagtc ccgaccctca gccgcatcca
     6721 cgtccatttt gaagacttca ctgctgagcg gagcggcggc gacgttggcc accttgacgg
     6781 cggcagccga gcaaatagcc cgctcctcgg ccagctgcga cactgccagg gattattccc
     6841 aaagcagcaa cagcagcagc aatagcaacg ctgccacagg agcaacagga actatggcca
     6901 gcggcaggac aggcagcagc aacagtttag aagagagcaa aagacttgag catcaacaac
     6961 aacaacaaca acagcagcaa caaccacaac agcatcagca gcaacaatat acatcaatac
     7021 tacacacggc cttaatgtca cagcgttcca tgccaccagc gacatcagct acatcagcag
     7081 cagcggcggt cacatcgacg ccgacaacgg cagcaacaac atcgcctacg cctgcgcatg
     7141 tgctgaaaac agcggtgacc ttagccagtc taagtgagat agccacagca cgtcagcagc
     7201 agcaggagca gcagcagcaa cagcagcagc agcagcagca gcaacaattg ctggcgatgc
     7261 aacagcaaca gcaacagcta acgaaactgc ccagaaggat aatcaacttt ggcagcaatc
     7321 acactgccaa cactgccacc aaagcattgg gagcgggatc agaagcagga gcaggagcag
     7381 gagtgggcat ggccacagca acggcaacag caacagttgg acgcaatcgc caactgggcc
     7441 tgagtcaggg tcaagtggtg aatggtaagc agaaagaatt gtgagccttg tgtttgcgat
     7501 aacgtttgtg tacccctcgt taacgctgcc agtcaggaca gcacgcctgc tcaatggcaa
     7561 ctgttgcggc cgattgcagg cgacagcagg aacaggggcc agcaggggca ccaacactgc
     7621 cggcagcaac acggtcaaca tgcagaccca gaccgacgat gagttgctgc cgctgttagc
     7681 catgctccca caagacgctc ccacaacaac aggagcagca gcaggagcag cagtatcatt
     7741 atcattatta gccacagcca cagccacaac cacatcttca tcatcatcat catcatcatc
     7801 attatcatca tcatcgacga tttcattagc aacacgtgca gcagcagtaa tgcagcagca
     7861 gcaacaatgg actgccgtgt cgacgacgcc agcaacagga gtgatgatga cgacgacgac
     7921 gacacaatca caatcacaag caacattagc accacaacaa cagcaactat cactgccaca
     7981 acaaaagcaa caatttaata cgagaataaa cagaaaaaca acaattacaa cagccaccac
     8041 aacagcacca acagcaacaa cagcaacaac aacaacaaca cgaaacagca acatcaactt
     8101 caacagcaac aataactata tgaataacaa taatataaac gccgccactg ccagagcctc
     8161 tgccttcgcc tccgcctctg ccatggccac ggccacgccc acaggcacac ccactgccac
     8221 gggcccgcgc cagagaccgg ccccgcaacc gcccccgccc ccgcgcacgc ccacagccac
     8281 ctcagcttct gttactgtag ttgtgttcaa aacaatgtta cctgatgact gaattatttt
     8341 agaatcgtac acatagagca tgttttatta tgattaccac tatgattacc acttacgatt
     8401 tacgattacg attatgatta cgattatgag tatgtctgga tatcattacc tattaactat
     8461 tcattagtta tggattatcg attattacta tttttactat tactattgct acatgcataa
     8521 tgtagagcct aattgatcta tcgtgtattt tagtttgtct aattgttgga tccctgtact
     8581 atatcccaca cacatacaca cactctcacc catatttgat cctcaccccc aaacaatcca
     8641 taagcacaca ctcatacaca cacacacaca cacacacaca ttgccgttac taaggaaggg
     8701 tatgcgcata tgcgaagttg aaacatttta atcatgtatt tttttttttt gtgtttttgc
     8761 tgaacggaga tgaaagttaa ggaggtcgag caacagcaat tagaagtgtt aattgagtag
     8821 gattatgagt acgtagcgag tatgtaacga gtatgtaatg actaagtaaa gtttacgtag
     8881 acacaatgcc acacacagct atagcaagca tttaggatgc gtgcatatgt cagttatcta
     8941 tttttttttt ttaaactaaa atctatataa agtaacgaga gaggaaggaa gagcgaagca
     9001 gggcgggaga ttcaaccaca taaagtataa atagtagagc ttgagccaag tagctagttt
     9061 agtgtacatc cgttttgggg cagggaagta gagcaatttg ttgacgagtc ggggtggtgg
     9121 ggtgagcggt aacgaagatc ggtaacggaa gtggagacaa agcggaaccg gaaacgggag
     9181 ctggactgcc agatcggaat cacacatcaa ccattatgag aacttcgctc gcaactcttg
     9241 cagaattata ctgttagatc ctgttagtta tccaacccaa aactcccaac ccacacacac
     9301 tatacctaag ttaagaacgc ataatcacat actacacata aattacgcat agtacataca
     9361 aaaaagatga agag