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Drosophila melanogaster hormone receptor 4, transcript variant F


LOCUS       NM_001272228            8417 bp    mRNA    linear   INV 26-DEC-2023
            (Hr4), mRNA.
ACCESSION   NM_001272228
VERSION     NM_001272228.1
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 8417)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 8417)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 8417)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 8417)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 8417)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 8417)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 8417)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 8417)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 8417)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 8417)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 8417)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 8417)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 8417)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 8417)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 8417)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 8417)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 8417)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 8417)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..8417
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..8417
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /exception="dicistronic gene"
                     /note="Hormone receptor 4"
                     /map="2B16-2C1"
                     /db_xref="FLYBASE:FBgn0264562"
                     /db_xref="GeneID:31162"
     CDS             660..5495
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="CG43934 gene product from transcript CG43934-RF;
                     CG43934-PF; Hr4-PF"
                     /codon_start=1
                     /product="hormone receptor 4, isoform F"
                     /protein_id="NP_001259157.1"
                     /db_xref="FLYBASE:FBpp0305530"
                     /db_xref="GeneID:31162"
                     /db_xref="FLYBASE:FBgn0264562"
                     /translation="MLPERVTIIPKIQTNTSTKSKFNEKSRTPHANSKNTHKSKCKLK
                     SKIGKKRKPPTIKTTTTTTIGKQENFLFLSKFEQSAKTRKNEIMTSIGIMTLSRGPYS
                     ELDKMSLFQDLKLKRRKIDSRCSSDGESIADTSTSSPDLLAPMSPKLCDSGSAGASLG
                     ASLPLPLALPLPMALPLPMSLPLPLTAASSAVTVSLAAVVAAVAETGGAGAGGAGTAV
                     TASGAGPCVSTSSTTAAAATSSTSSLSSSSSSSSSTSSSTSSASPTAGASSTATCPAS
                     SSSSSGNGSGGKSGSIKQEHTEIHSSSSAISAAAASTVMSPPPAEATRSSPATPEGGG
                     PAGDGSGATGGGNTSGGSTAGVAINEHQNNGNGSGGSSRASPDSLEEKPSTTTTTGRP
                     TLTPTNGVLSSASAGTGISTGSSAKLSEAGMSVIRSVKEERLLNVSSKMLVFHQQREQ
                     ETKAVAAAAAAAAAGHVTVLVTPSRIKSEPPPPASPSSTSSTQRERERERDRERDRER
                     ERERDRDREREREQSISSSQQHLSRVSASPPTQLSHGSLGPNIVQTHHLHQQLTQPLT
                     LRKSSPPTEHLLSQSMQHLTQQQAIHLHHLLGQQQQQQQASHPQQQQQQQHSPHSLVR
                     VKKEPNVGQRHLSPHHQQQSPLLQHHQQQQQQQQQQQQHLHQQQQQQQHHQQQPQALA
                     LMHPASLALRNSNRDAAILFRVKSEVHQQVAAGLPHLMQSAGGAAAAAAAAVAAQRMV
                     CFSNARINGVKPEVIGGPLGNLRPVGVGGGNGSGSVQCPSPHPSSSSSSSQLSPQTPS
                     QTPPRGTPTVIMGESCGVRTMVWGYEPPPPSAGQSHGQHPQQQQQSPHHQPQQQQQQQ
                     QQQSQQQQQQQQQQSLGQQQHCLSSPSAGSLTPSSSSGGGSVSGGGVGGPLTPSSVAP
                     QNNEEAAQLLLSLGQTRIQDMRSRPHPFRTPHALNMERLWAGDYSQLPPGQLQALNLS
                     AQQQQWGSSNSTGLGGVGGGMGGRNLEAPHEPTDEDEQPLVCMICEDKATGLHYGIIT
                     CEGCKGFFKRTVQNRRVYTCVADGTCEITKAQRNRCQYCRFKKCIEQGMVLQAVREDR
                     MPGGRNSGAVYNLYKVKYKKHKKTNQKQQQQAAQQQQQQAAAQQQHQQQQQHQQHQQH
                     QQQQLHSPLHHHHHQGHQSHHAQQQHHPQLSPHHLLSPQQQQLAAAVAAAAQHQQQQQ
                     QQQQQQQQAKLMGGVVDMKPMFLGPALKPELLQAPPMHSPAQQQQQQQQQQQQQQASP
                     HLSLSSPHQQQQQQQGQHQNHHQQQGGGGGGAGGGAQLPPHLVNGTILKTALTNPSEI
                     VHLRHRLDSAVSSSKDRQISYEHALGMIQTLIDCDAMEDIATLPHFSEFLEDKSEISE
                     KLCNIGDSIVHKLVSWTKKLPFYLEIPVEIHTKLLTDKWHEILILTTAAYQALHGKRR
                     GEGGGSRHGSPASTPLSTPTGTPLSTPIPSPAQPLHKDDPEFVSEVNSHLSTLQTCLT
                     TLMGQPIAMEQLKLDVGHMVDKMTQITIMFRRIKLKMEEYVCLKVYILLNKEVELESI
                     QERYVQVLRSYLQNSSPQNPQARLSELLSHIPEIQAAASLLLESKMFYVPFVLNSASI
                     R"
     misc_feature    3675..3944
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="DNA-binding domain of ecdysone-induced DHR4 orphan
                     nuclear receptor is composed of two C4-type zinc fingers;
                     Region: NR_DBD_DHR4_like; cd07168"
                     /db_xref="CDD:143542"
     misc_feature    order(3699..3701,3708..3710,3750..3752,3759..3761,
                     3807..3809,3825..3827,3855..3857,3864..3866)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="zinc binding site [ion binding]; other site"
                     /db_xref="CDD:143542"
     misc_feature    order(3729..3737,3753..3758,3762..3764,3768..3770,
                     3777..3779,3786..3788,3846..3851,3858..3860,3867..3869,
                     3900..3902,3909..3911)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative DNA binding site [nucleotide binding];
                     other site"
                     /db_xref="CDD:143542"
     misc_feature    4704..5480
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="The ligand binding domain of orphan nuclear
                     receptor Ecdysone-induced receptor DHR4; Region:
                     NR_LBD_DHR4_like; cd06953"
                     /db_xref="CDD:132751"
     misc_feature    order(4794..4799,4809..4811,4911..4913,4923..4925,
                     4932..4934,5130..5132,5199..5201)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative ligand binding site [chemical binding];
                     other site"
                     /db_xref="CDD:132751"
     misc_feature    order(4824..4826,4833..4835,4845..4847,4860..4862,
                     4875..4880,4884..4889,4896..4901)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative coactivator recognition site [active]"
                     /db_xref="CDD:132751"
ORIGIN      
        1 tttcacttta tttcgttcgc atgcacgcac gttggccgcg gagcaacatc tccagcaaac
       61 agaaatcgaa acctaaacag aaacagaaac agaagcaaaa aataaaacat aaaaaaaaaa
      121 caaaaaaaaa taaaaaacaa aacgaaacta caaatacaaa taacaaaaaa caaataacac
      181 ggcagcagcc gtcgctgaga cgcgaaactt ggctcgaact ttagtttagt ttttctcagt
      241 ttcagtggcc agtggctaaa gtgcggcgat atcgggcgaa acggttaacg gctcacggct
      301 catggctaac ggttgacagc cagcggttca tcacgcgcca aatccttttt tttcaagttt
      361 gtcaatgttt ttagtggtac taaagaaaac acatcattat atgtaaacta taggcaagaa
      421 tctaaagcca agagtgaatt gcagtcatcg aaaagcaacc ccaccccagt gcactaaact
      481 ggagcaacca acaattggcc aagaaaacaa acaagaacag aactcaccca gaaccaccca
      541 accaccaacc accaaccacc acccaccaaa aaccaaaaac cacccactgg aatgcgtact
      601 gccactgcca ctctgctctg cagctcaatg ggcagcgggc agggacgaac gcagtctaga
      661 tgcttcccga acgagtcaca ataataccaa aaatccaaac aaacacaagt acaaaatcca
      721 aattcaatga aaaatcaaga accccacatg caaactcaaa aaacacacac aaaagcaaat
      781 gcaaactgaa aagtaaaatc ggcaagaaaa gaaaaccccc aacaataaaa acaacaacaa
      841 caacaacgat cggcaagcaa gagaattttt tgtttttatc caaatttgaa caatcggcaa
      901 agacaagaaa aaacgaaatt atgacatcca taggaattat gacactgagc cgtggcccgt
      961 acagcgagct cgataaaatg agcctttttc aagacctcaa actcaaacgg cgcaaaatcg
     1021 attcgcgatg cagcagtgac ggcgagtcca tagcggacac gtccacctcg tcgccggacc
     1081 tgctggcgcc catgtcgccg aagctctgcg acagcggctc ggcgggggcg tcgctggggg
     1141 catcgctgcc cctgccgctg gccctgcccc tgccaatggc cctgccactg cccatgtcgc
     1201 tgcccctgcc cctcacggcg gcatcttcgg cggtcaccgt ttcgctggca gcggtcgtgg
     1261 ccgcggtggc cgagacgggt ggcgcgggcg cgggaggagc tgggacagca gtaacagcgt
     1321 cgggagcagg accatgcgtc tccacgtcgt ctacgacggc agcggcagcc acatcctcga
     1381 cctcctcgct ctcgtcctcc tcctcttcgt catcctccac gtcctccagc acttcctccg
     1441 cctcgccgac agctggagcc tcctccacgg ccacctgccc cgccagcagc agcagcagca
     1501 gtggaaacgg aagtgggggc aaaagtggta gcatcaagca ggagcacacg gagatacact
     1561 cgtcgagcag tgcgatttcg gcggccgccg cctcaacggt gatgtcaccg ccgcccgctg
     1621 aggcgacgag atccagtcca gccacgcccg agggaggcgg accagctggc gacggaagtg
     1681 gagcaacggg aggcggaaac acgagcggcg gatcaacggc tggagtggcc attaatgaac
     1741 accaaaacaa tggcaatggc agcggcggga gcagtcgagc ctctcccgat tcgctggaag
     1801 agaagccctc taccacaacg accacaggtc gtccaacgct cacgcccacg aatggggtgc
     1861 tgtcctccgc ctcggcgggc acggggattt ccacaggaag cagcgccaag ctgagcgagg
     1921 ctggtatgag tgtgatacgg tccgtgaagg aggagcgctt gctcaacgta tccagcaaga
     1981 tgctggtgtt ccatcagcag cgggagcaag agaccaaagc agtggcggct gcagcagcag
     2041 cagcagcggc gggccatgtg acggttctag tgacgccatc gcgcatcaaa tcggagccac
     2101 cgccgccggc ttcaccctcc tctacatcca gcacacaaag ggaaagggaa cgggaacgcg
     2161 atcgagagag ggatcgcgaa agggaacgcg agcgggaccg ggaccgggaa cgggaacggg
     2221 aacagtccat cagctcctcg cagcagcacc taagtcgggt ctccgccagt ccacccactc
     2281 agctgtccca cggcagcctg ggacccaaca ttgtgcagac gcaccatctt caccagcaac
     2341 tcacacagcc gctgacgctg cgcaagagca gcccgcccac agagcacctg ctcagtcagt
     2401 ccatgcaaca tctcacacag cagcaggcga tccacctgca tcacctactt ggccagcagc
     2461 agcagcagca gcaggcgtcg catccccagc agcaacagca gcagcaacac tcgccccact
     2521 ccctggtgcg ggtgaaaaag gaaccgaatg ttggtcagcg gcacttatcg ccgcatcacc
     2581 aacaacagtc gccactcctg cagcaccacc aacagcagca gcagcagcaa caacaacagc
     2641 aacagcatct gcatcagcaa cagcaacagc agcagcatca ccagcagcag ccccaggcac
     2701 tggccctgat gcatccggct tccctggcgc taaggaacag caatcgggat gcggccattc
     2761 tgtttcgggt gaagagcgaa gtgcaccagc aggtggccgc cgggctgccg catctgatgc
     2821 agtccgctgg tggggcagcg gccgccgccg cagcagctgt ggccgctcag cgaatggtat
     2881 gcttcagcaa tgccaggatc aatggcgtta agccggaggt gattggagga ccgctgggca
     2941 acctgcggcc cgtgggcgtc ggtggcggaa acggaagtgg ctccgtgcag tgcccctcgc
     3001 cgcatccatc ctcctcgtcg tcatcctcgc agctgtcgcc gcagacgccc tcccagacgc
     3061 cgccccgagg cacgcccacc gtcataatgg gcgagagctg cggggtgcgc accatggtct
     3121 ggggctacga gcctccgcca ccctcggcgg gccagtccca cggccagcac ccgcaacagc
     3181 aacagcagtc gccccaccac cagccgcaac aacaacagca gcagcaacaa cagcagtcgc
     3241 agcagcaaca gcaacagcag cagcaacagt cgctgggcca gcagcagcac tgcctctcct
     3301 cgccgtcggc gggatcgctg acgccctcct cttcgtccgg cggtggttcg gtatctggcg
     3361 gcggagtggg cggaccactc acaccctcct cggtggcgcc gcagaataac gaggaggccg
     3421 cccaactcct gctctccctg ggacagacac gcatccagga catgagatca cggccacacc
     3481 ccttccgcac accgcacgcc cttaatatgg agcggctgtg ggcgggagac tactcgcaat
     3541 tgccgcccgg ccagctgcag gctctgaatc tcagtgccca acagcagcag tggggcagca
     3601 gcaactccac gggtcttggt ggcgtaggcg gcggcatggg cggacgcaac ctggaggcgc
     3661 cgcacgagcc gaccgacgag gacgaacagc cgctcgtttg catgatctgc gaggacaagg
     3721 ccaccggcct gcactacggc atcatcacct gcgaggggtg caagggcttc ttcaagcgga
     3781 cggtgcagaa ccgacgagtc tacacctgcg tggcggacgg cacctgcgag ataaccaaag
     3841 cacagcgcaa ccgttgtcag tattgtcgat ttaagaagtg catcgagcag ggcatggtgc
     3901 tgcaagccgt tcgcgaggat cgcatgccgg gcggtcgcaa cagtggcgcc gtctacaatt
     3961 tgtacaaggt gaagtacaag aagcacaaga agaccaatca gaagcagcag cagcaggccg
     4021 cccagcagca gcagcagcag gcggcggcgc agcagcagca ccagcaacag cagcagcatc
     4081 aacagcacca gcaacatcag caacagcagt tgcactcgcc gctccaccat caccaccacc
     4141 agggccacca gtcgcaccac gcgcagcagc agcaccaccc acagctgtcg ccgcaccacc
     4201 tgctgtcgcc gcagcagcag caacttgccg ccgcggtggc agcagctgcg cagcaccaac
     4261 agcaacagca acaacagcag caacagcagc agcaggccaa gctgatgggc ggcgtggtgg
     4321 acatgaagcc catgttcctc ggccccgctt tgaagccgga gttgctgcaa gcacccccca
     4381 tgcacagtcc ggcccagcaa caacaacagc agcagcagca gcagcagcaa cagcaggcct
     4441 cgccgcatct ctcgcttagc tcaccgcacc agcagcagca gcagcagcag ggacagcacc
     4501 aaaaccacca ccagcaacaa ggtgggggtg gcggaggagc tggtggagga gctcaactgc
     4561 cgccgcacct ggtgaacgga acgatactga agacggccct aaccaatccc agcgagattg
     4621 tacatctgcg ccaccgcctc gactcggcgg tcagttcgtc caaggaccga cagatctcgt
     4681 acgagcacgc cttaggcatg atccagacac tgatcgactg cgacgcgatg gaggacatag
     4741 ccacactgcc gcacttcagc gagttccttg aggacaagtc ggagattagc gagaaactgt
     4801 gcaacatcgg cgattccata gtccacaagc tggtgtcgtg gacaaaaaag ttgcccttct
     4861 acctggagat cccggtggag atacatacca aactactgac ggacaagtgg cacgagatcc
     4921 ttatcctgac cacggccgcc taccaggcgt tgcatggcaa gcggcgtggc gagggaggag
     4981 gcagcaggca tggttcgccg gcgtcaacgc cactgagcac gcccactggt acgccgttga
     5041 gcacaccgat accctcgccc gcccagccac tgcacaagga cgacccggag tttgtcagcg
     5101 aggtgaactc gcacctgagc acactgcaaa cctgcttgac cacgctaatg ggccagccga
     5161 tagcgatgga gcagctgaag ctggacgtcg ggcacatggt ggacaagatg acccagatca
     5221 ccatcatgtt ccggcgaatc aagctcaaga tggaggagta cgtctgcctg aaggtttaca
     5281 tactgctaaa caaagaagtg gaactggaga gcatccagga gcggtacgtc caggtgctgc
     5341 gctcctacct gcaaaactcc tcgccgcaga atccgcaggc gaggctcagt gaactgctct
     5401 cccacatacc agagatccag gctgcggcta gcctgctgct cgagagcaag atgttctatg
     5461 tgcccttcgt gctcaactcg gcgagcataa ggtagcagat gcttgagcac ggactgctgc
     5521 ccaccgccaa ccatcagtcg gcgccaccac ctggtcctga tacgatcccg gagcggcagc
     5581 agcaacagca acaggagcgg cgtcgaagca ggcagcagca gcaccagttg ctgcgcaacc
     5641 tgcccaagat caagatcgag atcggttccc agcgcgatcc ggaggaagag gagtcggagc
     5701 aagagctaga gcaggagccg gagcaagagc aggatgtgca gtcccgaccc tcagccgcat
     5761 ccacgtccat tttgaagact tcactgctga gcggagcggc ggcgacgttg gccaccttga
     5821 cggcggcagc cgagcaaata gcccgctcct cggccagctg cgacactgcc agggattatt
     5881 cccaaagcag caacagcagc agcaatagca acgctgccac aggagcaaca ggaactatgg
     5941 ccagcggcag gacaggcagc agcaacagtt tagaagagag caaaagactt gagcatcaac
     6001 aacaacaaca acaacagcag caacaaccac aacagcatca gcagcaacaa tatacatcaa
     6061 tactacacac ggccttaatg tcacagcgtt ccatgccacc agcgacatca gctacatcag
     6121 cagcagcggc ggtcacatcg acgccgacaa cggcagcaac aacatcgcct acgcctgcgc
     6181 atgtgctgaa aacagcggtg accttagcca gtctaagtga gatagccaca gcacgtcagc
     6241 agcagcagga gcagcagcag caacagcagc agcagcagca gcagcaacaa ttgctggcga
     6301 tgcaacagca acagcaacag ctaacgaaac tgcccagaag gataatcaac tttggcagca
     6361 atcacactgc caacactgcc accaaagcat tgggagcggg atcagaagca ggagcaggag
     6421 caggagtggg catggccaca gcaacggcaa cagcaacagt tggacgcaat cgccaactgg
     6481 gcctgagtca gggtcaagtg gtgaatggta agcagaaaga attgtgagcc ttgtgtttgc
     6541 gataacgttt gtgtacccct cgttaacgct gccagtcagg acagcacgcc tgctcaatgg
     6601 caactgttgc ggccgattgc aggcgacagc aggaacaggg gccagcaggg gcaccaacac
     6661 tgccggcagc aacacggtca acatgcagac ccagaccgac gatgagttgc tgccgctgtt
     6721 agccatgctc ccacaagacg ctcccacaac aacaggagca gcagcaggag cagcagtatc
     6781 attatcatta ttagccacag ccacagccac aaccacatct tcatcatcat catcatcatc
     6841 atcattatca tcatcatcga cgatttcatt agcaacacgt gcagcagcag taatgcagca
     6901 gcagcaacaa tggactgccg tgtcgacgac gccagcaaca ggagtgatga tgacgacgac
     6961 gacgacacaa tcacaatcac aagcaacatt agcaccacaa caacagcaac tatcactgcc
     7021 acaacaaaag caacaattta atacgagaat aaacagaaaa acaacaatta caacagccac
     7081 cacaacagca ccaacagcaa caacagcaac aacaacaaca acacgaaaca gcaacatcaa
     7141 cttcaacagc aacaataact atatgaataa caataatata aacgccgcca ctgccagagc
     7201 ctctgccttc gcctccgcct ctgccatggc cacggccacg cccacaggca cacccactgc
     7261 cacgggcccg cgccagagac cggccccgca accgcccccg cccccgcgca cgcccacagc
     7321 cacctcagct tctgttactg tagttgtgtt caaaacaatg ttacctgatg actgaattat
     7381 tttagaatcg tacacataga gcatgtttta ttatgattac cactatgatt accacttacg
     7441 atttacgatt acgattatga ttacgattat gagtatgtct ggatatcatt acctattaac
     7501 tattcattag ttatggatta tcgattatta ctatttttac tattactatt gctacatgca
     7561 taatgtagag cctaattgat ctatcgtgta ttttagtttg tctaattgtt ggatccctgt
     7621 actatatccc acacacatac acacactctc acccatattt gatcctcacc cccaaacaat
     7681 ccataagcac acactcatac acacacacac acacacacac acattgccgt tactaaggaa
     7741 gggtatgcgc atatgcgaag ttgaaacatt ttaatcatgt attttttttt tttgtgtttt
     7801 tgctgaacgg agatgaaagt taaggaggtc gagcaacagc aattagaagt gttaattgag
     7861 taggattatg agtacgtagc gagtatgtaa cgagtatgta atgactaagt aaagtttacg
     7921 tagacacaat gccacacaca gctatagcaa gcatttagga tgcgtgcata tgtcagttat
     7981 ctattttttt tttttaaact aaaatctata taaagtaacg agagaggaag gaagagcgaa
     8041 gcagggcggg agattcaacc acataaagta taaatagtag agcttgagcc aagtagctag
     8101 tttagtgtac atccgttttg gggcagggaa gtagagcaat ttgttgacga gtcggggtgg
     8161 tggggtgagc ggtaacgaag atcggtaacg gaagtggaga caaagcggaa ccggaaacgg
     8221 gagctggact gccagatcgg aatcacacat caaccattat gagaacttcg ctcgcaactc
     8281 ttgcagaatt atactgttag atcctgttag ttatccaacc caaaactccc aacccacaca
     8341 cactatacct aagttaagaa cgcataatca catactacac ataaattacg catagtacat
     8401 acaaaaaaga tgaagag