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Drosophila melanogaster hormone receptor 4, transcript variant G


LOCUS       NM_001272227            9386 bp    mRNA    linear   INV 26-DEC-2023
            (Hr4), mRNA.
ACCESSION   NM_001272227
VERSION     NM_001272227.1
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 9386)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 9386)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 9386)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 9386)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 9386)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 9386)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 9386)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 9386)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 9386)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 9386)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 9386)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 9386)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 9386)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 9386)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 9386)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 9386)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 9386)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 9386)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..9386
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..9386
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /exception="dicistronic gene"
                     /note="Hormone receptor 4"
                     /map="2B16-2C1"
                     /db_xref="FLYBASE:FBgn0264562"
                     /db_xref="GeneID:31162"
     CDS             504..6464
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="CG43934 gene product from transcript CG43934-RG;
                     CG43934-PG; Hr4-PG"
                     /codon_start=1
                     /product="hormone receptor 4, isoform G"
                     /protein_id="NP_001259156.1"
                     /db_xref="FLYBASE:FBpp0305529"
                     /db_xref="GeneID:31162"
                     /db_xref="FLYBASE:FBgn0264562"
                     /translation="MTPLQRDQLNHQQFVHHYLQSQPVNNKIDSFDIKHIRKTQNPTT
                     NPKFRTEDRFKTSSKSKITNPESGNTQLHFHLSETPDLSASTSLLPSAFVASSAALVN
                     VTCQAEEGQSSAGSHYTVHRGNDAGISFAALDFRFIRRAAQMNAQQPLILEPGSKASS
                     VVAAAAELEAATTTAALATSSATEASAADSYPAVQQNGGHEEQRLNGNHGCRREQPDD
                     RTDQPEQLDQQDEPEPVKDVANKLTIIKTPLNKTLLKKCNSNGSLQMPHANNGSSINI
                     NNRNNTNNTNNNNSNNNNNAASNDVCDSGLQSGNSNEQEPFANNDRHHHIYHHHHHHY
                     HHHHGGDKSEASGDVASSSSADPNSQVKDEPTAQDSQDNQAAGAAEAGGATAVAATCS
                     PSKCNKCNSHNNNSNCNNNNNTISNNNNNTSSNTNTNSNSNNNANNNNNNNNNNHYNN
                     YNNYYKKKSRMPPICRIMTLSRGPYSELDKMSLFQDLKLKRRKIDSRCSSDGESIADT
                     STSSPDLLAPMSPKLCDSGSAGASLGASLPLPLALPLPMALPLPMSLPLPLTAASSAV
                     TVSLAAVVAAVAETGGAGAGGAGTAVTASGAGPCVSTSSTTAAAATSSTSSLSSSSSS
                     SSSTSSSTSSASPTAGASSTATCPASSSSSSGNGSGGKSGSIKQEHTEIHSSSSAISA
                     AAASTVMSPPPAEATRSSPATPEGGGPAGDGSGATGGGNTSGGSTAGVAINEHQNNGN
                     GSGGSSRASPDSLEEKPSTTTTTGRPTLTPTNGVLSSASAGTGISTGSSAKLSEAGMS
                     VIRSVKEERLLNVSSKMLVFHQQREQETKAVAAAAAAAAAGHVTVLVTPSRIKSEPPP
                     PASPSSTSSTQRERERERDRERDRERERERDRDREREREQSISSSQQHLSRVSASPPT
                     QLSHGSLGPNIVQTHHLHQQLTQPLTLRKSSPPTEHLLSQSMQHLTQQQAIHLHHLLG
                     QQQQQQQASHPQQQQQQQHSPHSLVRVKKEPNVGQRHLSPHHQQQSPLLQHHQQQQQQ
                     QQQQQQHLHQQQQQQQHHQQQPQALALMHPASLALRNSNRDAAILFRVKSEVHQQVAA
                     GLPHLMQSAGGAAAAAAAAVAAQRMVCFSNARINGVKPEVIGGPLGNLRPVGVGGGNG
                     SGSVQCPSPHPSSSSSSSQLSPQTPSQTPPRGTPTVIMGESCGVRTMVWGYEPPPPSA
                     GQSHGQHPQQQQQSPHHQPQQQQQQQQQQSQQQQQQQQQQSLGQQQHCLSSPSAGSLT
                     PSSSSGGGSVSGGGVGGPLTPSSVAPQNNEEAAQLLLSLGQTRIQDMRSRPHPFRTPH
                     ALNMERLWAGDYSQLPPGQLQALNLSAQQQQWGSSNSTGLGGVGGGMGGRNLEAPHEP
                     TDEDEQPLVCMICEDKATGLHYGIITCEGCKGFFKRTVQNRRVYTCVADGTCEITKAQ
                     RNRCQYCRFKKCIEQGMVLQAVREDRMPGGRNSGAVYNLYKVKYKKHKKTNQKQQQQA
                     AQQQQQQAAAQQQHQQQQQHQQHQQHQQQQLHSPLHHHHHQGHQSHHAQQQHHPQLSP
                     HHLLSPQQQQLAAAVAAAAQHQQQQQQQQQQQQQAKLMGGVVDMKPMFLGPALKPELL
                     QAPPMHSPAQQQQQQQQQQQQQQASPHLSLSSPHQQQQQQQGQHQNHHQQQGGGGGGA
                     GGGAQLPPHLVNGTILKTALTNPSEIVHLRHRLDSAVSSSKDRQISYEHALGMIQTLI
                     DCDAMEDIATLPHFSEFLEDKSEISEKLCNIGDSIVHKLVSWTKKLPFYLEIPVEIHT
                     KLLTDKWHEILILTTAAYQALHGKRRGEGGGSRHGSPASTPLSTPTGTPLSTPIPSPA
                     QPLHKDDPEFVSEVNSHLSTLQTCLTTLMGQPIAMEQLKLDVGHMVDKMTQITIMFRR
                     IKLKMEEYVCLKVYILLNKAEVELESIQERYVQVLRSYLQNSSPQNPQARLSELLSHI
                     PEIQAAASLLLESKMFYVPFVLNSASIR"
     misc_feature    4641..4910
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="DNA-binding domain of ecdysone-induced DHR4 orphan
                     nuclear receptor is composed of two C4-type zinc fingers;
                     Region: NR_DBD_DHR4_like; cd07168"
                     /db_xref="CDD:143542"
     misc_feature    order(4665..4667,4674..4676,4716..4718,4725..4727,
                     4773..4775,4791..4793,4821..4823,4830..4832)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="zinc binding site [ion binding]; other site"
                     /db_xref="CDD:143542"
     misc_feature    order(4695..4703,4719..4724,4728..4730,4734..4736,
                     4743..4745,4752..4754,4812..4817,4824..4826,4833..4835,
                     4866..4868,4875..4877)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative DNA binding site [nucleotide binding];
                     other site"
                     /db_xref="CDD:143542"
     misc_feature    5670..6449
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /old_locus_tag="Dmel_CG16902"
                     /old_locus_tag="Dmel_CG3600"
                     /old_locus_tag="Dmel_CG42527"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="The ligand binding domain of orphan nuclear
                     receptor Ecdysone-induced receptor DHR4; Region:
                     NR_LBD_DHR4_like; cd06953"
                     /db_xref="CDD:132751"
     misc_feature    order(5760..5765,5775..5777,5877..5879,5889..5891,
                     5898..5900,6096..6098,6165..6167)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative ligand binding site [chemical binding];
                     other site"
                     /db_xref="CDD:132751"
     misc_feature    order(5790..5792,5799..5801,5811..5813,5826..5828,
                     5841..5846,5850..5855,5862..5867)
                     /gene="Hr4"
                     /locus_tag="Dmel_CG43934"
                     /gene_synonym="CG16902; CG3600; CG42527; CG43692; CG43934;
                     DHR4; Dmel\CG43934; EG:133E12.2; EG:BACH61I5.1; EP(X)1232;
                     EP1232; GRF; hr4; HR4; NR6A2; null"
                     /note="putative coactivator recognition site [active]"
                     /db_xref="CDD:132751"
ORIGIN      
        1 gtgtatccca cgagctagtc gcgcgcggaa agcaaacgct tcgaaaatca aacgatattt
       61 ccgttttgac gtttaacccc cacccaccca aataaagcaa atctaactaa agagtcctgc
      121 aaaaaaatca acagctgctt gatcttgaag cctccatatc ttgaaatcct gcatcagcca
      181 acagccccaa aacaaaacaa aacaaaacga atgcaaatac actaatgtga acaacaacta
      241 aattgctagt gaaatgtttt aagccaaatg ctgtgagtgt gtaacaaaaa aaaaaaaaaa
      301 aaaaaaaaaa aaagaagagt gattcaaagc aaccgcaagc aatcaacttg taacggtagt
      361 cggatactcg cttgcccgcg gagcatggat atcgccccag ttccgccacc accattatcg
      421 atatccgtat cgctttcggc tccaggatca acggtattcg caaacgctgg ccactagaga
      481 agcgtcagcc gaatgcctca aagatgacac cgctgcaacg ggaccaattg aaccaccaac
      541 aatttgtgca ccactatttg caaagtcaac ctgtcaataa caaaattgac agctttgata
      601 ttaaacatat acgtaaaacc caaaacccga ccacaaaccc aaagtttcga actgaagacc
      661 gtttcaagac aagcagcaag agcaagatta ccaacccgga gagcggaaat acccaactcc
      721 attttcattt gagtgaaacg ccagacttat ccgcctcaac ttcgctgttg ccgtcagcat
      781 ttgttgcatc ctccgccgcc ttggtgaatg tgacatgcca ggcggaggaa ggacaaagtt
      841 cagcaggatc acactacaca gttcatagag ggaatgacgc aggcatttct tttgcagcgt
      901 tggactttcg ttttataaga agagctgccc agatgaatgc ccagcagccg ctgatcctgg
      961 agcctggatc caaggccagc agtgttgttg cggcagctgc cgagctagag gcagcaacaa
     1021 ccacggcagc gctggccacg tcctccgcca cggaggcttc tgcagcggac agctatcctg
     1081 ccgtacagca gaacggcggc cacgaggagc agcggctcaa tggcaaccac ggatgtcgcc
     1141 gcgagcagcc ggatgatcgg acggaccagc cggagcagtt ggatcagcag gatgagcccg
     1201 agccagtcaa ggacgtggcc aacaagctga ccattatcaa gacgccgttg aacaagactc
     1261 tgctgaagaa gtgcaacagc aacgggagcc tgcagatgcc gcacgccaac aatggcagca
     1321 gcatcaacat caataatcgc aacaacacaa acaacaccaa caacaacaat agcaacaata
     1381 acaacaatgc ggccagcaac gacgtctgcg attccggttt gcaaagcgga aatagcaatg
     1441 aacaagaacc atttgccaac aacgatcgcc atcaccacat ctaccaccac caccaccatc
     1501 actatcatca tcaccatggc ggcgataaga gcgaggcgag cggcgacgtt gcatcctcgt
     1561 cctccgcgga ccccaactcg caggtcaagg acgagcccac agcgcaggat agccaggaca
     1621 atcaagcggc cggagctgca gaggctggag gagctactgc ggttgcagcc acgtgcagcc
     1681 ccagcaagtg caacaagtgc aactcgcaca ataacaacag caactgcaac aataacaata
     1741 ataccatcag caacaacaac aataatacaa gcagtaatac caatacgaac agcaacagca
     1801 acaacaatgc caacaacaat aataataata acaataataa ccattacaac aactacaata
     1861 actattataa gaaaaagtct cgtatgccgc ccatttgccg aattatgaca ctgagccgtg
     1921 gcccgtacag cgagctcgat aaaatgagcc tttttcaaga cctcaaactc aaacggcgca
     1981 aaatcgattc gcgatgcagc agtgacggcg agtccatagc ggacacgtcc acctcgtcgc
     2041 cggacctgct ggcgcccatg tcgccgaagc tctgcgacag cggctcggcg ggggcgtcgc
     2101 tgggggcatc gctgcccctg ccgctggccc tgcccctgcc aatggccctg ccactgccca
     2161 tgtcgctgcc cctgcccctc acggcggcat cttcggcggt caccgtttcg ctggcagcgg
     2221 tcgtggccgc ggtggccgag acgggtggcg cgggcgcggg aggagctggg acagcagtaa
     2281 cagcgtcggg agcaggacca tgcgtctcca cgtcgtctac gacggcagcg gcagccacat
     2341 cctcgacctc ctcgctctcg tcctcctcct cttcgtcatc ctccacgtcc tccagcactt
     2401 cctccgcctc gccgacagct ggagcctcct ccacggccac ctgccccgcc agcagcagca
     2461 gcagcagtgg aaacggaagt gggggcaaaa gtggtagcat caagcaggag cacacggaga
     2521 tacactcgtc gagcagtgcg atttcggcgg ccgccgcctc aacggtgatg tcaccgccgc
     2581 ccgctgaggc gacgagatcc agtccagcca cgcccgaggg aggcggacca gctggcgacg
     2641 gaagtggagc aacgggaggc ggaaacacga gcggcggatc aacggctgga gtggccatta
     2701 atgaacacca aaacaatggc aatggcagcg gcgggagcag tcgagcctct cccgattcgc
     2761 tggaagagaa gccctctacc acaacgacca caggtcgtcc aacgctcacg cccacgaatg
     2821 gggtgctgtc ctccgcctcg gcgggcacgg ggatttccac aggaagcagc gccaagctga
     2881 gcgaggctgg tatgagtgtg atacggtccg tgaaggagga gcgcttgctc aacgtatcca
     2941 gcaagatgct ggtgttccat cagcagcggg agcaagagac caaagcagtg gcggctgcag
     3001 cagcagcagc agcggcgggc catgtgacgg ttctagtgac gccatcgcgc atcaaatcgg
     3061 agccaccgcc gccggcttca ccctcctcta catccagcac acaaagggaa agggaacggg
     3121 aacgcgatcg agagagggat cgcgaaaggg aacgcgagcg ggaccgggac cgggaacggg
     3181 aacgggaaca gtccatcagc tcctcgcagc agcacctaag tcgggtctcc gccagtccac
     3241 ccactcagct gtcccacggc agcctgggac ccaacattgt gcagacgcac catcttcacc
     3301 agcaactcac acagccgctg acgctgcgca agagcagccc gcccacagag cacctgctca
     3361 gtcagtccat gcaacatctc acacagcagc aggcgatcca cctgcatcac ctacttggcc
     3421 agcagcagca gcagcagcag gcgtcgcatc cccagcagca acagcagcag caacactcgc
     3481 cccactccct ggtgcgggtg aaaaaggaac cgaatgttgg tcagcggcac ttatcgccgc
     3541 atcaccaaca acagtcgcca ctcctgcagc accaccaaca gcagcagcag cagcaacaac
     3601 aacagcaaca gcatctgcat cagcaacagc aacagcagca gcatcaccag cagcagcccc
     3661 aggcactggc cctgatgcat ccggcttccc tggcgctaag gaacagcaat cgggatgcgg
     3721 ccattctgtt tcgggtgaag agcgaagtgc accagcaggt ggccgccggg ctgccgcatc
     3781 tgatgcagtc cgctggtggg gcagcggccg ccgccgcagc agctgtggcc gctcagcgaa
     3841 tggtatgctt cagcaatgcc aggatcaatg gcgttaagcc ggaggtgatt ggaggaccgc
     3901 tgggcaacct gcggcccgtg ggcgtcggtg gcggaaacgg aagtggctcc gtgcagtgcc
     3961 cctcgccgca tccatcctcc tcgtcgtcat cctcgcagct gtcgccgcag acgccctccc
     4021 agacgccgcc ccgaggcacg cccaccgtca taatgggcga gagctgcggg gtgcgcacca
     4081 tggtctgggg ctacgagcct ccgccaccct cggcgggcca gtcccacggc cagcacccgc
     4141 aacagcaaca gcagtcgccc caccaccagc cgcaacaaca acagcagcag caacaacagc
     4201 agtcgcagca gcaacagcaa cagcagcagc aacagtcgct gggccagcag cagcactgcc
     4261 tctcctcgcc gtcggcggga tcgctgacgc cctcctcttc gtccggcggt ggttcggtat
     4321 ctggcggcgg agtgggcgga ccactcacac cctcctcggt ggcgccgcag aataacgagg
     4381 aggccgccca actcctgctc tccctgggac agacacgcat ccaggacatg agatcacggc
     4441 cacacccctt ccgcacaccg cacgccctta atatggagcg gctgtgggcg ggagactact
     4501 cgcaattgcc gcccggccag ctgcaggctc tgaatctcag tgcccaacag cagcagtggg
     4561 gcagcagcaa ctccacgggt cttggtggcg taggcggcgg catgggcgga cgcaacctgg
     4621 aggcgccgca cgagccgacc gacgaggacg aacagccgct cgtttgcatg atctgcgagg
     4681 acaaggccac cggcctgcac tacggcatca tcacctgcga ggggtgcaag ggcttcttca
     4741 agcggacggt gcagaaccga cgagtctaca cctgcgtggc ggacggcacc tgcgagataa
     4801 ccaaagcaca gcgcaaccgt tgtcagtatt gtcgatttaa gaagtgcatc gagcagggca
     4861 tggtgctgca agccgttcgc gaggatcgca tgccgggcgg tcgcaacagt ggcgccgtct
     4921 acaatttgta caaggtgaag tacaagaagc acaagaagac caatcagaag cagcagcagc
     4981 aggccgccca gcagcagcag cagcaggcgg cggcgcagca gcagcaccag caacagcagc
     5041 agcatcaaca gcaccagcaa catcagcaac agcagttgca ctcgccgctc caccatcacc
     5101 accaccaggg ccaccagtcg caccacgcgc agcagcagca ccacccacag ctgtcgccgc
     5161 accacctgct gtcgccgcag cagcagcaac ttgccgccgc ggtggcagca gctgcgcagc
     5221 accaacagca acagcaacaa cagcagcaac agcagcagca ggccaagctg atgggcggcg
     5281 tggtggacat gaagcccatg ttcctcggcc ccgctttgaa gccggagttg ctgcaagcac
     5341 cccccatgca cagtccggcc cagcaacaac aacagcagca gcagcagcag cagcaacagc
     5401 aggcctcgcc gcatctctcg cttagctcac cgcaccagca gcagcagcag cagcagggac
     5461 agcaccaaaa ccaccaccag caacaaggtg ggggtggcgg aggagctggt ggaggagctc
     5521 aactgccgcc gcacctggtg aacggaacga tactgaagac ggccctaacc aatcccagcg
     5581 agattgtaca tctgcgccac cgcctcgact cggcggtcag ttcgtccaag gaccgacaga
     5641 tctcgtacga gcacgcctta ggcatgatcc agacactgat cgactgcgac gcgatggagg
     5701 acatagccac actgccgcac ttcagcgagt tccttgagga caagtcggag attagcgaga
     5761 aactgtgcaa catcggcgat tccatagtcc acaagctggt gtcgtggaca aaaaagttgc
     5821 ccttctacct ggagatcccg gtggagatac ataccaaact actgacggac aagtggcacg
     5881 agatccttat cctgaccacg gccgcctacc aggcgttgca tggcaagcgg cgtggcgagg
     5941 gaggaggcag caggcatggt tcgccggcgt caacgccact gagcacgccc actggtacgc
     6001 cgttgagcac accgataccc tcgcccgccc agccactgca caaggacgac ccggagtttg
     6061 tcagcgaggt gaactcgcac ctgagcacac tgcaaacctg cttgaccacg ctaatgggcc
     6121 agccgatagc gatggagcag ctgaagctgg acgtcgggca catggtggac aagatgaccc
     6181 agatcaccat catgttccgg cgaatcaagc tcaagatgga ggagtacgtc tgcctgaagg
     6241 tttacatact gctaaacaaa gcagaagtgg aactggagag catccaggag cggtacgtcc
     6301 aggtgctgcg ctcctacctg caaaactcct cgccgcagaa tccgcaggcg aggctcagtg
     6361 aactgctctc ccacatacca gagatccagg ctgcggctag cctgctgctc gagagcaaga
     6421 tgttctatgt gcccttcgtg ctcaactcgg cgagcataag gtagcagatg cttgagcacg
     6481 gactgctgcc caccgccaac catcagtcgg cgccaccacc tggtcctgat acgatcccgg
     6541 agcggcagca gcaacagcaa caggagcggc gtcgaagcag gcagcagcag caccagttgc
     6601 tgcgcaacct gcccaagatc aagatcgaga tcggttccca gcgcgatccg gaggaagagg
     6661 agtcggagca agagctagag caggagccgg agcaagagca ggatgtgcag tcccgaccct
     6721 cagccgcatc cacgtccatt ttgaagactt cactgctgag cggagcggcg gcgacgttgg
     6781 ccaccttgac ggcggcagcc gagcaaatag cccgctcctc ggccagctgc gacactgcca
     6841 gggattattc ccaaagcagc aacagcagca gcaatagcaa cgctgccaca ggagcaacag
     6901 gaactatggc cagcggcagg acaggcagca gcaacagttt agaagagagc aaaagacttg
     6961 agcatcaaca acaacaacaa caacagcagc aacaaccaca acagcatcag cagcaacaat
     7021 atacatcaat actacacacg gccttaatgt cacagcgttc catgccacca gcgacatcag
     7081 ctacatcagc agcagcggcg gtcacatcga cgccgacaac ggcagcaaca acatcgccta
     7141 cgcctgcgca tgtgctgaaa acagcggtga ccttagccag tctaagtgag atagccacag
     7201 cacgtcagca gcagcaggag cagcagcagc aacagcagca gcagcagcag cagcaacaat
     7261 tgctggcgat gcaacagcaa cagcaacagc taacgaaact gcccagaagg ataatcaact
     7321 ttggcagcaa tcacactgcc aacactgcca ccaaagcatt gggagcggga tcagaagcag
     7381 gagcaggagc aggagtgggc atggccacag caacggcaac agcaacagtt ggacgcaatc
     7441 gccaactggg cctgagtcag ggtcaagtgg tgaatggtaa gcagaaagaa ttgtgagcct
     7501 tgtgtttgcg ataacgtttg tgtacccctc gttaacgctg ccagtcagga cagcacgcct
     7561 gctcaatggc aactgttgcg gccgattgca ggcgacagca ggaacagggg ccagcagggg
     7621 caccaacact gccggcagca acacggtcaa catgcagacc cagaccgacg atgagttgct
     7681 gccgctgtta gccatgctcc cacaagacgc tcccacaaca acaggagcag cagcaggagc
     7741 agcagtatca ttatcattat tagccacagc cacagccaca accacatctt catcatcatc
     7801 atcatcatca tcattatcat catcatcgac gatttcatta gcaacacgtg cagcagcagt
     7861 aatgcagcag cagcaacaat ggactgccgt gtcgacgacg ccagcaacag gagtgatgat
     7921 gacgacgacg acgacacaat cacaatcaca agcaacatta gcaccacaac aacagcaact
     7981 atcactgcca caacaaaagc aacaatttaa tacgagaata aacagaaaaa caacaattac
     8041 aacagccacc acaacagcac caacagcaac aacagcaaca acaacaacaa cacgaaacag
     8101 caacatcaac ttcaacagca acaataacta tatgaataac aataatataa acgccgccac
     8161 tgccagagcc tctgccttcg cctccgcctc tgccatggcc acggccacgc ccacaggcac
     8221 acccactgcc acgggcccgc gccagagacc ggccccgcaa ccgcccccgc ccccgcgcac
     8281 gcccacagcc acctcagctt ctgttactgt agttgtgttc aaaacaatgt tacctgatga
     8341 ctgaattatt ttagaatcgt acacatagag catgttttat tatgattacc actatgatta
     8401 ccacttacga tttacgatta cgattatgat tacgattatg agtatgtctg gatatcatta
     8461 cctattaact attcattagt tatggattat cgattattac tatttttact attactattg
     8521 ctacatgcat aatgtagagc ctaattgatc tatcgtgtat tttagtttgt ctaattgttg
     8581 gatccctgta ctatatccca cacacataca cacactctca cccatatttg atcctcaccc
     8641 ccaaacaatc cataagcaca cactcataca cacacacaca cacacacaca cattgccgtt
     8701 actaaggaag ggtatgcgca tatgcgaagt tgaaacattt taatcatgta tttttttttt
     8761 ttgtgttttt gctgaacgga gatgaaagtt aaggaggtcg agcaacagca attagaagtg
     8821 ttaattgagt aggattatga gtacgtagcg agtatgtaac gagtatgtaa tgactaagta
     8881 aagtttacgt agacacaatg ccacacacag ctatagcaag catttaggat gcgtgcatat
     8941 gtcagttatc tatttttttt ttttaaacta aaatctatat aaagtaacga gagaggaagg
     9001 aagagcgaag cagggcggga gattcaacca cataaagtat aaatagtaga gcttgagcca
     9061 agtagctagt ttagtgtaca tccgttttgg ggcagggaag tagagcaatt tgttgacgag
     9121 tcggggtggt ggggtgagcg gtaacgaaga tcggtaacgg aagtggagac aaagcggaac
     9181 cggaaacggg agctggactg ccagatcgga atcacacatc aaccattatg agaacttcgc
     9241 tcgcaactct tgcagaatta tactgttaga tcctgttagt tatccaaccc aaaactccca
     9301 acccacacac actataccta agttaagaac gcataatcac atactacaca taaattacgc
     9361 atagtacata caaaaaagat gaagag