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Drosophila melanogaster Tousled-like kinase (Tlk), transcript


LOCUS       NM_001144680            6513 bp    mRNA    linear   INV 26-DEC-2023
            variant F, mRNA.
ACCESSION   NM_001144680
VERSION     NM_001144680.3
DBLINK      BioProject: PRJNA164
            BioSample: SAMN02803731
KEYWORDS    RefSeq.
SOURCE      Drosophila melanogaster (fruit fly)
  ORGANISM  Drosophila melanogaster
            Eukaryota; Metazoa; Ecdysozoa; Arthropoda; Hexapoda; Insecta;
            Pterygota; Neoptera; Endopterygota; Diptera; Brachycera;
            Muscomorpha; Ephydroidea; Drosophilidae; Drosophila; Sophophora.
REFERENCE   1  (bases 1 to 6513)
  AUTHORS   Matthews,B.B., Dos Santos,G., Crosby,M.A., Emmert,D.B., St
            Pierre,S.E., Gramates,L.S., Zhou,P., Schroeder,A.J., Falls,K.,
            Strelets,V., Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: Impact of
            High-Throughput Data
  JOURNAL   G3 (Bethesda) 5 (8), 1721-1736 (2015)
   PUBMED   26109357
  REMARK    Publication Status: Online-Only
REFERENCE   2  (bases 1 to 6513)
  AUTHORS   Crosby,M.A., Gramates,L.S., Dos Santos,G., Matthews,B.B., St
            Pierre,S.E., Zhou,P., Schroeder,A.J., Falls,K., Emmert,D.B.,
            Russo,S.M. and Gelbart,W.M.
  CONSRTM   FlyBase Consortium
  TITLE     Gene Model Annotations for Drosophila melanogaster: The
            Rule-Benders
  JOURNAL   G3 (Bethesda) 5 (8), 1737-1749 (2015)
   PUBMED   26109356
  REMARK    Publication Status: Online-Only
REFERENCE   3  (bases 1 to 6513)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Wan,K.H., Park,S., Mendez,I.,
            Galle,S.E., Booth,B.W., Pfeiffer,B.D., George,R.A., Svirskas,R.,
            Krzywinski,M., Schein,J., Accardo,M.C., Damia,E., Messina,G.,
            Mendez-Lago,M., de Pablos,B., Demakova,O.V., Andreyeva,E.N.,
            Boldyreva,L.V., Marra,M., Carvalho,A.B., Dimitri,P., Villasante,A.,
            Zhimulev,I.F., Rubin,G.M., Karpen,G.H. and Celniker,S.E.
  TITLE     The Release 6 reference sequence of the Drosophila melanogaster
            genome
  JOURNAL   Genome Res 25 (3), 445-458 (2015)
   PUBMED   25589440
REFERENCE   4  (bases 1 to 6513)
  AUTHORS   Hoskins,R.A., Carlson,J.W., Kennedy,C., Acevedo,D., Evans-Holm,M.,
            Frise,E., Wan,K.H., Park,S., Mendez-Lago,M., Rossi,F.,
            Villasante,A., Dimitri,P., Karpen,G.H. and Celniker,S.E.
  TITLE     Sequence finishing and mapping of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1625-1628 (2007)
   PUBMED   17569867
REFERENCE   5  (bases 1 to 6513)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  TITLE     The Release 5.1 annotation of Drosophila melanogaster
            heterochromatin
  JOURNAL   Science 316 (5831), 1586-1591 (2007)
   PUBMED   17569856
  REMARK    Erratum:[Science. 2007 Sep 7;317(5843):1325]
REFERENCE   6  (bases 1 to 6513)
  AUTHORS   Quesneville,H., Bergman,C.M., Andrieu,O., Autard,D., Nouaud,D.,
            Ashburner,M. and Anxolabehere,D.
  TITLE     Combined evidence annotation of transposable elements in genome
            sequences
  JOURNAL   PLoS Comput Biol 1 (2), 166-175 (2005)
   PUBMED   16110336
REFERENCE   7  (bases 1 to 6513)
  AUTHORS   Hoskins,R.A., Smith,C.D., Carlson,J.W., Carvalho,A.B., Halpern,A.,
            Kaminker,J.S., Kennedy,C., Mungall,C.J., Sullivan,B.A.,
            Sutton,G.G., Yasuhara,J.C., Wakimoto,B.T., Myers,E.W.,
            Celniker,S.E., Rubin,G.M. and Karpen,G.H.
  TITLE     Heterochromatic sequences in a Drosophila whole-genome shotgun
            assembly
  JOURNAL   Genome Biol 3 (12), RESEARCH0085 (2002)
   PUBMED   12537574
REFERENCE   8  (bases 1 to 6513)
  AUTHORS   Kaminker,J.S., Bergman,C.M., Kronmiller,B., Carlson,J.,
            Svirskas,R., Patel,S., Frise,E., Wheeler,D.A., Lewis,S.E.,
            Rubin,G.M., Ashburner,M. and Celniker,S.E.
  TITLE     The transposable elements of the Drosophila melanogaster
            euchromatin: a genomics perspective
  JOURNAL   Genome Biol 3 (12), RESEARCH0084 (2002)
   PUBMED   12537573
REFERENCE   9  (bases 1 to 6513)
  AUTHORS   Misra,S., Crosby,M.A., Mungall,C.J., Matthews,B.B., Campbell,K.S.,
            Hradecky,P., Huang,Y., Kaminker,J.S., Millburn,G.H., Prochnik,S.E.,
            Smith,C.D., Tupy,J.L., Whitfied,E.J., Bayraktaroglu,L.,
            Berman,B.P., Bettencourt,B.R., Celniker,S.E., de Grey,A.D.,
            Drysdale,R.A., Harris,N.L., Richter,J., Russo,S., Schroeder,A.J.,
            Shu,S.Q., Stapleton,M., Yamada,C., Ashburner,M., Gelbart,W.M.,
            Rubin,G.M. and Lewis,S.E.
  TITLE     Annotation of the Drosophila melanogaster euchromatic genome: a
            systematic review
  JOURNAL   Genome Biol 3 (12), RESEARCH0083 (2002)
   PUBMED   12537572
REFERENCE   10 (bases 1 to 6513)
  AUTHORS   Celniker,S.E., Wheeler,D.A., Kronmiller,B., Carlson,J.W.,
            Halpern,A., Patel,S., Adams,M., Champe,M., Dugan,S.P., Frise,E.,
            Hodgson,A., George,R.A., Hoskins,R.A., Laverty,T., Muzny,D.M.,
            Nelson,C.R., Pacleb,J.M., Park,S., Pfeiffer,B.D., Richards,S.,
            Sodergren,E.J., Svirskas,R., Tabor,P.E., Wan,K., Stapleton,M.,
            Sutton,G.G., Venter,C., Weinstock,G., Scherer,S.E., Myers,E.W.,
            Gibbs,R.A. and Rubin,G.M.
  TITLE     Finishing a whole-genome shotgun: release 3 of the Drosophila
            melanogaster euchromatic genome sequence
  JOURNAL   Genome Biol 3 (12), RESEARCH0079 (2002)
   PUBMED   12537568
REFERENCE   11 (bases 1 to 6513)
  AUTHORS   Adams,M.D., Celniker,S.E., Holt,R.A., Evans,C.A., Gocayne,J.D.,
            Amanatides,P.G., Scherer,S.E., Li,P.W., Hoskins,R.A., Galle,R.F.,
            George,R.A., Lewis,S.E., Richards,S., Ashburner,M., Henderson,S.N.,
            Sutton,G.G., Wortman,J.R., Yandell,M.D., Zhang,Q., Chen,L.X.,
            Brandon,R.C., Rogers,Y.H., Blazej,R.G., Champe,M., Pfeiffer,B.D.,
            Wan,K.H., Doyle,C., Baxter,E.G., Helt,G., Nelson,C.R., Gabor,G.L.,
            Abril,J.F., Agbayani,A., An,H.J., Andrews-Pfannkoch,C., Baldwin,D.,
            Ballew,R.M., Basu,A., Baxendale,J., Bayraktaroglu,L., Beasley,E.M.,
            Beeson,K.Y., Benos,P.V., Berman,B.P., Bhandari,D., Bolshakov,S.,
            Borkova,D., Botchan,M.R., Bouck,J., Brokstein,P., Brottier,P.,
            Burtis,K.C., Busam,D.A., Butler,H., Cadieu,E., Center,A.,
            Chandra,I., Cherry,J.M., Cawley,S., Dahlke,C., Davenport,L.B.,
            Davies,P., de Pablos,B., Delcher,A., Deng,Z., Mays,A.D., Dew,I.,
            Dietz,S.M., Dodson,K., Doup,L.E., Downes,M., Dugan-Rocha,S.,
            Dunkov,B.C., Dunn,P., Durbin,K.J., Evangelista,C.C., Ferraz,C.,
            Ferriera,S., Fleischmann,W., Fosler,C., Gabrielian,A.E., Garg,N.S.,
            Gelbart,W.M., Glasser,K., Glodek,A., Gong,F., Gorrell,J.H., Gu,Z.,
            Guan,P., Harris,M., Harris,N.L., Harvey,D., Heiman,T.J.,
            Hernandez,J.R., Houck,J., Hostin,D., Houston,K.A., Howland,T.J.,
            Wei,M.H., Ibegwam,C., Jalali,M., Kalush,F., Karpen,G.H., Ke,Z.,
            Kennison,J.A., Ketchum,K.A., Kimmel,B.E., Kodira,C.D., Kraft,C.,
            Kravitz,S., Kulp,D., Lai,Z., Lasko,P., Lei,Y., Levitsky,A.A.,
            Li,J., Li,Z., Liang,Y., Lin,X., Liu,X., Mattei,B., McIntosh,T.C.,
            McLeod,M.P., McPherson,D., Merkulov,G., Milshina,N.V., Mobarry,C.,
            Morris,J., Moshrefi,A., Mount,S.M., Moy,M., Murphy,B., Murphy,L.,
            Muzny,D.M., Nelson,D.L., Nelson,D.R., Nelson,K.A., Nixon,K.,
            Nusskern,D.R., Pacleb,J.M., Palazzolo,M., Pittman,G.S., Pan,S.,
            Pollard,J., Puri,V., Reese,M.G., Reinert,K., Remington,K.,
            Saunders,R.D., Scheeler,F., Shen,H., Shue,B.C., Siden-Kiamos,I.,
            Simpson,M., Skupski,M.P., Smith,T., Spier,E., Spradling,A.C.,
            Stapleton,M., Strong,R., Sun,E., Svirskas,R., Tector,C., Turner,R.,
            Venter,E., Wang,A.H., Wang,X., Wang,Z.Y., Wassarman,D.A.,
            Weinstock,G.M., Weissenbach,J., Williams,S.M., WoodageT,
            Worley,K.C., Wu,D., Yang,S., Yao,Q.A., Ye,J., Yeh,R.F.,
            Zaveri,J.S., Zhan,M., Zhang,G., Zhao,Q., Zheng,L., Zheng,X.H.,
            Zhong,F.N., Zhong,W., Zhou,X., Zhu,S., Zhu,X., Smith,H.O.,
            Gibbs,R.A., Myers,E.W., Rubin,G.M. and Venter,J.C.
  TITLE     The genome sequence of Drosophila melanogaster
  JOURNAL   Science 287 (5461), 2185-2195 (2000)
   PUBMED   10731132
REFERENCE   12 (bases 1 to 6513)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Pfeiffer,B., Frise,E., George,R.,
            Hoskins,R., Stapleton,M., Pacleb,J., Park,S., Svirskas,R.,
            Smith,E., Yu,C. and Rubin,G.
  CONSRTM   Berkeley Drosophila Genome Project
  TITLE     Drosophila melanogaster release 4 sequence
  JOURNAL   Unpublished
REFERENCE   13 (bases 1 to 6513)
  CONSRTM   NCBI Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (20-DEC-2023) National Center for Biotechnology
            Information, NIH, Bethesda, MD 20894, USA
REFERENCE   14 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (13-DEC-2023) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   15 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (19-OCT-2022) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   16 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (20-APR-2020) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   17 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (22-APR-2019) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   18 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (24-MAY-2018) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   19 (bases 1 to 6513)
  CONSRTM   FlyBase
  TITLE     Direct Submission
  JOURNAL   Submitted (07-DEC-2016) FlyBase, Harvard University, Biological
            Laboratories, 16 Divinity Ave, Cambridge, MA 02138, USA
REFERENCE   20 (bases 1 to 6513)
  AUTHORS   Celniker,S., Carlson,J., Kennedy,C., Wan,K., Frise,E., Hoskins,R.,
            Park,S., Svirskas,R. and Karpen,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One #Cyclotron RoadOne
            Cyclotron Road, MS 64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   21 (bases 1 to 6513)
  AUTHORS   Celniker,S., Carlson,J., Wan,K., Frise,E., Hoskins,R., Park,S.,
            Svirskas,R. and Rubin,G.
  TITLE     Direct Submission
  JOURNAL   Submitted (10-AUG-2006) Berkeley Drosophila Genome Project,
            Lawrence Berkeley National Laboratory, One Cyclotron Road, MS
            64-121, Berkeley, CA 94720, USA
  REMARK    Direct Submission
REFERENCE   22 (bases 1 to 6513)
  AUTHORS   Smith,C.D., Shu,S., Mungall,C.J. and Karpen,G.H.
  CONSRTM   Drosophila Heterochromatin Genome Project
  TITLE     Direct Submission
  JOURNAL   Submitted (01-AUG-2006) Drosophila Heterochromatin Genome Project,
            Ernest Orlando Lawrence Berkeley National Laboratory, 1 Cyclotron
            Road, Mailstop 64-121, Berkeley, CA 94720, USA
REFERENCE   23 (bases 1 to 6513)
  AUTHORS   Adams,M.D., Celniker,S.E., Gibbs,R.A., Rubin,G.M. and Venter,C.J.
  TITLE     Direct Submission
  JOURNAL   Submitted (21-MAR-2000) Celera Genomics, 45 West Gude Drive,
            Rockville, MD 20850, USA
COMMENT     REVIEWED REFSEQ: This record has been curated by FlyBase. This
            record is derived from an annotated genomic sequence (NC_004354).
            
            On Jan 16, 2013 this sequence version replaced NM_001144680.2.
            
            ##Genome-Annotation-Data-START##
            Annotation Provider :: FlyBase
            Annotation Status   :: Full annotation
            Annotation Version  :: Release 6.54
            URL                 :: http://flybase.org
            ##Genome-Annotation-Data-END##
FEATURES             Location/Qualifiers
     source          1..6513
                     /organism="Drosophila melanogaster"
                     /mol_type="mRNA"
                     /db_xref="taxon:7227"
                     /chromosome="X"
                     /genotype="y[1]; Gr22b[1] Gr22d[1] cn[1] CG33964[R4.2]
                     bw[1] sp[1]; LysC[1] MstProx[1] GstD5[1] Rh6[1]"
     gene            1..6513
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Tousled-like kinase"
                     /map="3E6-3F1"
                     /db_xref="FLYBASE:FBgn0283657"
                     /db_xref="GeneID:318206"
     CDS             299..4099
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /EC_number="2.7.11.1"
                     /note="CG34412 gene product from transcript CG34412-RF;
                     CG34412-PF; Tlk-PF; BEST:GH17330; tousled-like kinase;
                     tousled like kinase"
                     /codon_start=1
                     /product="Tousled-like kinase, isoform F"
                     /protein_id="NP_001138152.1"
                     /db_xref="FLYBASE:FBpp0288855"
                     /db_xref="GeneID:318206"
                     /db_xref="FLYBASE:FBgn0283657"
                     /translation="MDQFQTSLNPRKLELLESRITGVRMSPGAHLQMSPQNTSSLSQH
                     HPHQQQQLQPPQQQQQHFPNHHSAQQQSQQQQQQEQQNPQQQAQQQQQILPHQHLQHL
                     HKHPHQLQLHQQQQQQLHQQQQQHFHQQSLQGLHQGSSNPDSNMSTGSSHSEKDVNDM
                     LSGGAATPGAAAAAIQQQHPAFAPTLGMQQPPPPPPQHSNNGGEMGYLSAGTTTTTSV
                     LTVGKPRTPAERKRKRKMPPCATSADEAGSGGGSGGAGATVVNNSSLKGKSLAFRDMP
                     KVNMSLNLGDRLGGSAGSGVGAGGAGSGGGGAGSGSGSGGGKSARLMLPVSDNKKIND
                     YFNKQQTGVGVGVPGGAGGNTAGLRGSHTGGGSKSPSSAQQQQTAAQQQGSGVATGGS
                     AGGSAGNQVQVQTSSAYALYPPASPQTQTSQQQQQQQPGSDFHYVNSSKAQQQQQRQQ
                     QQTSNQMVPPHVVVGLGGHPLSLASIQQQTPLSQQQQQQQQQQQQQQLGPPTTSTASV
                     VPTHPHQLGSLGVVGMVGVGVGVGVGVNVGVGPPLPPPPPMAMPAAIITYSKATQTEV
                     SLHELQEREAEHESGKVKLDEMTRLSDEQKSQIVGNQKTIDQHKCHIAKCIDVVKKLL
                     KEKSSIEKKEARQKCMQNRLRLGQFVTQRVGATFQENWTDGYAFQELSRRQEEITAER
                     EEIDRQKKQLMKKRPAESGRKRNNNSNQNNQQQQQQQHQQQQQQQNSNSNDSTQLTSG
                     VVTGPGSDRVSVSVDSGLGGNNAGAIGGGTVGGGVGGGGVGGGGVGGGGGRGLSRSNS
                     TQANQAQLLHNGGGGSGGNVGNSGGVGDRLSDRGGGGGGIGGNDSGSCSDSGTFLKPD
                     PVSGAYTAQEYYEYDEILKLRQNALKKEDADLQLEMEKLERERNLHIRELKRILNEDQ
                     SRFNNHPVLNDRYLLLMLLGKGGFSEVHKAFDLKEQRYVACKVHQLNKDWKEDKKANY
                     IKHALREYNIHKALDHPRVVKLYDVFEIDANSFCTVLEYCDGHDLDFYLKQHKTIPER
                     EARSIIMQVVSALKYLNEIKPPVIHYDLKPGNILLTEGNVCGEIKITDFGLSKVMDDE
                     NYNPDHGMDLTSQGAGTYWYLPPECFVVGKNPPKISSKVDVWSVGVIFYQCLYGKKPF
                     GHNQSQATILEENTILKATEVQFSNKPTVSNEAKSFIRGCLAYRKEDRMDVFALARHE
                     YIQPPIPKHGRGSLNQQQQAQQQQQQQQQQQQQQSSTSQANSTGQTSFSAHMFGNMNQ
                     SSSS"
     misc_feature    <1976..>2386
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Family of unknown function (DUF5401); Region:
                     DUF5401; pfam17380"
                     /db_xref="CDD:375164"
     misc_feature    3110..3913
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /old_locus_tag="CG32781"
                     /old_locus_tag="CG32782"
                     /old_locus_tag="Dmel_CG32781"
                     /old_locus_tag="Dmel_CG32782"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="Catalytic domain of the Serine/Threonine kinase,
                     Tousled-Like Kinase; Region: STKc_TLK; cd13990"
                     /db_xref="CDD:270892"
     misc_feature    order(3119..3121,3158..3160,3164..3166,3272..3274,
                     3323..3334,3344..3346,3350..3352,3467..3469,3473..3475,
                     3479..3484,3488..3490,3530..3532,3539..3541,3608..3619)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="active site"
                     /db_xref="CDD:270892"
     misc_feature    order(3344..3346,3350..3352,3467..3469,3473..3475,
                     3479..3481,3539..3541,3608..3619)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="polypeptide substrate binding site [polypeptide
                     binding]; other site"
                     /db_xref="CDD:270892"
     misc_feature    order(3527..3565,3578..3619)
                     /gene="Tlk"
                     /locus_tag="Dmel_CG34412"
                     /gene_synonym="anon-WO03040301.114; anon-WO03040301.116;
                     BcDNA:GH07910; BEST:GH17330; CG12462; CG2829; CG32781;
                     CG32782; CG33219; CG34412; Dmel\CG34412; dtlk; EP(X)1408;
                     EP(X)1413; EP1408; EP1413; GH17330; l(1)G0054; tlk; TLK"
                     /note="activation loop (A-loop); other site"
                     /db_xref="CDD:270892"
ORIGIN      
        1 atatttttag gccatttctg cgtgtgtgcg ttcgtgtgaa tattgtgaat atttttttta
       61 ctgtgaaaat agacaaaaat cggctgtgga aaaccgaaat gaatttgtca ctttattatt
      121 gccatggctc tgacaaatct tgtcgcactc gttgccgaca tttgcaatcg atcaatatta
      181 aaatattgaa taactaaacc aaccacgagc taaaagaaaa atttcagaaa aacttttgta
      241 tctgaatcga aaacaaaaaa aaaaacaaat tcctatcggc gcttataatc acaaagtaat
      301 ggatcaattt caaacttcac taaatccgcg gaaactcgag cttctggagt ctcgaattac
      361 tggagtgcgg atgtcccccg gcgcccattt gcagatgtcc ccgcagaaca cttcgtccct
      421 aagtcaacac catccacatc aacagcaaca gttacaaccc ccacagcagc aacaacagca
      481 tttccctaac catcacagcg cccagcaaca gtcgcagcag cagcagcaac aggagcaaca
      541 gaatccccag cagcaggcgc aacagcagca gcagatactc ccacatcaac atttgcagca
      601 cctgcacaag catccgcatc agctgcaact gcatcagcag cagcaacaac aactccacca
      661 gcaacagcag caacacttcc accagcagtc gctgcaaggg ctgcatcagg gtagcagcaa
      721 tccggattcg aatatgagca ctggctcctc gcatagcgag aaggatgtca atgatatgct
      781 gagtggcggt gcagcaacgc caggagctgc agcagcagcg attcaacagc aacatcccgc
      841 ctttgcgccc acactgggaa tgcagcaacc accgccgccc ccacctcaac actccaataa
      901 tggaggcgag atgggctact tgtcggcagg cacgaccacg acgacgtcgg tgttaacggt
      961 aggcaagcct cggacgccag cggagcggaa acggaagcga aaaatgcctc catgtgccac
     1021 tagtgcggat gaggcgggga gtggcggtgg ctctggcgga gcaggagcaa ccgttgttaa
     1081 caacagcagc ctgaagggca aatcattggc ctttcgtgat atgcccaagg taaacatgag
     1141 cctgaatctg ggcgatcgtc tgggaggatc tgcaggaagc ggagtaggag ccggtggcgc
     1201 cggaagcggg ggaggtggcg ctggttccgg ttctggaagc ggtggcggca aaagcgcccg
     1261 cctgatgctg ccagtcagcg acaacaagaa gatcaacgac tatttcaata agcagcaaac
     1321 gggcgtgggc gtcggtgtgc caggtggtgc gggaggcaat accgctggcc ttcgaggatc
     1381 acatacggga ggtggcagca agtcaccctc atccgcccag cagcagcaaa cggcggcaca
     1441 gcagcaggga agcggtgttg cgacgggagg cagtgcaggc ggttccgctg gcaaccaggt
     1501 gcaagtgcaa acgagcagcg cttacgccct ttacccacca gctagtcccc aaacccagac
     1561 gtcacagcaa cagcagcagc agcaaccggg atcagacttt cactatgtca actccagcaa
     1621 ggcgcagcaa caacagcagc gtcaacagca acagacttcc aatcaaatgg ttcctccaca
     1681 cgtggtcgtt ggccttggtg gtcatccact gagcctcgcg tccattcagc agcagacgcc
     1741 cttatcccag cagcaacagc agcaacaaca gcagcagcaa cagcagcaac tgggaccacc
     1801 gaccacatcg acggcctccg tcgtgccaac gcatccgcat caactcggat ccctgggagt
     1861 tgttgggatg gtcggtgtgg gtgttggcgt gggcgttgga gtaaatgtgg gtgtgggacc
     1921 accactgcca ccaccaccgc cgatggccat gccagcggcc attatcactt atagtaaggc
     1981 cactcaaacg gaggtgtcgc tgcatgaatt gcaggagcgc gaagcggagc acgaatcggg
     2041 caaggtgaag ctagacgaga tgacacggct gtccgatgaa caaaagtccc aaattgttgg
     2101 caaccagaag acgattgacc agcacaagtg ccacatagcc aagtgtattg atgtggtcaa
     2161 gaagctgttg aaggagaaga gcagcatcga gaagaaggag gcgcgacaga agtgcatgca
     2221 gaatcgcctc aggctcggac agtttgttac ccaacgagtg ggcgccacat tccaggagaa
     2281 ctggacggac ggctatgcgt tccaggagct gagtcggcgg caagaagaaa taaccgctga
     2341 gcgtgaagag atagatcggc agaaaaagca gctgatgaaa aagcgtccgg cggagtccgg
     2401 acgcaagcgc aacaacaaca gtaaccagaa caaccagcag cagcagcaac agcaacacca
     2461 gcaacagcag cagcaacaaa attccaactc gaacgattcc acgcagctga cgagcggagt
     2521 tgttaccggt ccaggcagtg atcgtgtgag cgtaagcgtc gacagcggat tgggtggcaa
     2581 taatgcgggc gcgatcggtg gcggaaccgt tggtggtggc gttggaggtg gtggtgttgg
     2641 aggcggtggt gtcggaggcg gcggtggacg tggactttct cgcagcaatt cgacgcaggc
     2701 caatcaggct caattgctgc acaacggcgg tggtggttcg ggcggcaatg tcggcaactc
     2761 gggcggcgtt ggcgaccgct tgtcagatcg aggaggagga ggtggcggca tcggcggaaa
     2821 cgatagcggc agctgctcgg actcgggcac tttcctgaag ccagaccccg tatcgggtgc
     2881 ctacacagcg caggagtatt acgagtacga tgagatcctc aagttgcgac aaaatgccct
     2941 caaaaaggag gacgccgacc tgcagctgga gatggagaag ctggagcggg agcgcaatct
     3001 gcacatccga gagctcaagc ggattcttaa cgaggatcag tcccgcttta acaatcatcc
     3061 cgtgctgaat gatcgctatc ttctgttgat gctcctgggc aagggcggct tctcagaggt
     3121 ccacaaggcc ttcgacctga aggagcaacg ctatgtcgca tgtaaggtgc accaattaaa
     3181 caaggattgg aaggaggata agaaagctaa ttatatcaaa cacgctttgc gggaatacaa
     3241 cattcacaag gcactggatc atccgcgggt cgtcaagcta tacgatgtct tcgagatcga
     3301 tgcgaattcc ttttgcacag tgctcgaata ctgtgatggc cacgatctgg acttctattt
     3361 gaagcaacat aagactatac ccgagcgtga agcgcgctcg ataataatgc aggttgtatc
     3421 tgcactcaag tatctaaatg agattaagcc tccagttatc cactacgatc tgaagcccgg
     3481 caacattctg cttaccgagg gcaacgtctg cggcgagatt aagatcaccg acttcggtct
     3541 gtcaaaggtg atggacgacg agaattacaa tcccgatcac ggcatggatc tgacctctca
     3601 gggggcggga acctactggt atctgccacc cgagtgcttt gtcgtgggca aaaatccgcc
     3661 gaaaatctcc tccaaagtgg acgtatggag tgtgggtgtt atcttctacc agtgtctgta
     3721 cggcaaaaag cccttcggtc acaatcagtc gcaggccacg attctcgagg agaatacgat
     3781 cctgaaggcc accgaagtgc agttctccaa caagccaacc gtttctaacg aggccaagag
     3841 tttcattcgg ggatgcttgg cctatcgcaa ggaggatcgc atggatgtgt tcgcactggc
     3901 caggcacgag tacattcagc caccgatacc gaaacatggg cgcggttcgc tcaatcagca
     3961 acagcaggcg caacaacagc agcagcaaca acagcaacag cagcagcaac agtcgtcgac
     4021 gtcacaggcc aattctacag gccagacatc tttctctgcc cacatgtttg gcaatatgaa
     4081 tcagtcgagt tcgtcctagc tgccaaccaa gcgcaattcg aactcataat tcgccgccgc
     4141 cgcagtagca gcagctacat tagctgccag tagtcaacag caacttcaac accagcaaca
     4201 gcaactccaa atccagcaat accagcagaa tcagcaactt catttgcaaa acagcagtag
     4261 caattatgcg gccacgcttc tagatgcagc ggccttttgc tacgaaaccg tgatggatgc
     4321 agcccacttc caaaacaaca ataataacag caccatcatc aagaacaata tgcagacaat
     4381 gccggcaaac gctgccaatg ctgctgcagc tgcaggggga gggggaacac cgacctatca
     4441 ggggaaatac aagcgctggc agcagcaaca gttacaacag caggcgcaac tccaacagca
     4501 gcagcacaag ctgctgttgc agcaagagct gctctaaaat atgaagggaa tgatgacaag
     4561 cgagtttaag agaacgagga gctgcctgcc acactgccgc cgtcagcaat aacaacatcc
     4621 caagaaaatt cacagcgagt cgcaaaagca tcaacaaaca aagcaaaaaa cgcaaaaaag
     4681 gattacaaag acgcagagaa taggggacat ttgcaatggg aaataacaaa ttattgcatt
     4741 atctgctgga ttgtgattgt ggttcaaaca ttaaaaatac acttactcat acacacaagg
     4801 tagacactgc agactaattt tgccttaaaa gaaacaaaac gaataatgtt aactgtaaat
     4861 tttcactttg tgtccatgca tagttactag taatgatgat ataaatatat aaatatatat
     4921 aaaatataaa aattatataa atgtataaat ttatatttat atatatatat atatatatat
     4981 atatatatct gcataaacat tgaagcatac tagtaaatct ttataaatat atttatatat
     5041 atatttatct cttcctttta acgcttttaa tcggaaaaaa catcccaatt atattttaat
     5101 tcaatttgaa atataaaagt catttttgtt tgttttgtat caaaatccaa aaaaaattga
     5161 tttaaatata aaaataccat aaacattaac accacacaca cacacacaca cacatatata
     5221 tatatatata tatatatata tatatataca tatatatata catttgtata tagtcgtatc
     5281 tgaaccacaa acaaatccac agataaaaaa aactgaaaaa ttcatagaat caaaccattt
     5341 ttttatatgc cacaaagaaa agaaaataag tagaaatgca agaaaaatcg agaaaaagcg
     5401 taattcatga gatcagatcg gatcagaaaa atcaacagaa aataaaccta tttgaagtaa
     5461 aatacttcta aatacatagt aaactaatgt aatagttttt attaaaatta cacatatatg
     5521 tatcgatata tatacatgtt tttttttttt taattttaaa attataattg taatgcacta
     5581 tgcgtaaatt gtattaaata ttaatatcca aggcatatta atagaaatat atatacatat
     5641 ataaaacctg caaaaaaaaa acatgaacta ccaagctaaa tgtttcaata atatatagat
     5701 ctatatgtat gtaaaaccga tatgaaatct gcctacagcc agacaaaaaa aaccttaaag
     5761 agaaagatca gaagaggaac agaagagtat gagcggggag tgaaagatgc aagattcatc
     5821 caaaagcgaa aaacattttt ttcttaagtc tacgcaaaac aaaatgaatg aaaagtaacc
     5881 cgaaattagt ttttagagta taagtataaa aaatggaaaa tatcgcgata gaaagaaaca
     5941 aaaatggacg cagataagga cggcaagcag aacaagagta ataaagcagc ggcaagaact
     6001 ccctaaaaat agcaaagagc attgtaaata tttatataaa taatttatgc ctaattaaag
     6061 tttaaattaa tgaagaaaac cagcaaaaaa acataaaaaa ccagaagaac catttcagat
     6121 ttgtatgcaa atttgtatat gtacgtactt tcctttaatt ttagccaatc agtttgtaag
     6181 attaaatttt ttttttgcat ctctataaag aataatgata tgaggataat gattccgatt
     6241 tttttttccg tttaatgtaa ccgttagaat tagcccataa tggtcgatat ctaaatgtgt
     6301 gtgtatatcg atgtgatgtc atcgattgtc gataacgatt ggatgcgagt gtgtgttaat
     6361 ttaatttgat aaaagataaa ggaaagcaag gaatgtgaag ataatgaaag gagatgcata
     6421 gagaatgaga gcgaatctat ctattatata attatgtaaa atgcacatct ttaaattata
     6481 aattatacat atttaaaagg aaactattat tat